Korean ctenostome bryozoans-observations on living colonies, new records, five new species, and an updated checklist
Author
Seo, Ji-Eun
Author
Chae, Hyun Sook
Author
Winston, Judith E.
Author
Zágoršek, Kamil
Author
Gordon, Dennis P.
text
Zootaxa
2018
2018-09-28
4486
3
251
283
journal article
29193
10.11646/zootaxa.4486.3.3
8bf7deb9-6336-4232-aa5f-cf6805630d2a
1175-5326
1453233
B87F5447-A747-4D96-8845-0B30B40412A3
Nolella
cf.
papuensis
(
Busk, 1886
)
(
Fig. 8
)
Cylindroecium papuense
Busk, 1886
: 38
, pl. 8, fig. 2.
Nolella papuensis
:
Harmer 1915
: 53
, pl. 4, figs 10–20;
Rho & Kim 1981
: 61
, pl. 4, figs 1, 2;
Rho & Seo 1986
: 33
;
Seo 2005
: 283
, pl. 14, fig. 23
Material examined.
MBRBK
1705
,
9
May
2017,
Munseom Island
offshore of
Seogwipo Harbor
,
Jeju
Island,
20 m
.
Description.
Colony forming chain or network of narrow cylindrical zooids. Zooid tubes varying in length, straight to slightly curved, connected by very narrow stoloniform elongations; zooids brown, opaque, their cuticle covered by particles of sediment proximally (
Fig. 8A, B
), the distal ends of zooids usually lighter and more translucent (
Fig. 8D
), with fine annular rings and topped distally by the often-squared orifice (
Fig. 8E, F
). Mean zooid length
1.043 mm
(range
0.97–1.09 mm
, N = 3); zooid width, mean
0.186 mm
(range 0.145–0.217, N = 3), no dilation in diameter at base. Tentacle crown equitentacled, tentacles thin, straightly held; the one tentacle crown that expanded in our study (
Fig. 8C
) had 12 tentacles. No evidence of reproduction in colonies found.
Remarks.
The muddy-looking cylindrical zooids of
Nolella
resemble tubes of polychaetes or amphipods, but their squared-off orifice is diagnostic, distinguishing them as belonging to
Nolella
. The zooids of the specimens collected in this study are shorter than the average for zooids of
N. stipata
recorded in Florida (2.1 x
1.8 mm
zooid length and width) (
Winston 1982
) and
Brazil
(2.3 x
1.8 mm
) (
Vieira
et al
. 2014
). Korean material collected previously and identified as
Nolella papuensis
had zooids 1.0–
2.8 mm
long and
0.20–0.30 mm
wide (
Seo 2005
).
Harmer’s (1915)
account of
Nolella papuen
sis from the
Siboga Expedition
reported considerable variability in zooid length depending on state of growth and/or regeneration, but able to attain almost
4 mm
in height with a tentacle crown of c. 18 tentacles.
Busk’s (1886)
original description gave much smaller dimensions (1.3 x
0.10–0.11 mm
). We believe the Korean specimens (all three from the South Sea), most closely approach
N. papuensis
. Embryos were lacking in our material, but embryos in colonies from other areas are brooded, several in a row, in the distal half of zooid tubes.
Distribution.
Indonesia
, New
Guinea
.
Korea
: Goheung, South Sea; Seogwipo,
Jeju
Island; intertidal to
20 m
.