Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae)
Author
Yasunaga, Tomohide
Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA;
Author
Tamada, Yui
Nagasaki West High School, Biology Club, Takenokubo 12 -
Author
Hinami, Haruka
Nagasaki West High School, Biology Club, Takenokubo 12 -
Author
Miyazaki, Ayana
Nagasaki West High School, Biology Club, Takenokubo 12 -
Author
Duwal, Ram Keshari
Visiting Researcher, Agriculture & Agri-Food Canada Environmental Health, K. W. Neatby: Bldg # 20, 960 Carling Avenue, Central Experimental Farm, Ottawa Ontario, Canada K
Author
Nagashima, Tetsuya
Nagasaki West High School, Biology Club, Takenokubo 12 -
text
Acta Entomologica Musei Nationalis Pragae
2019
Acta. Ent. Mus. Natl. Pragae
2019-02-26
59
1
71
99
http://dx.doi.org/10.2478/aemnp-2019-0007
journal article
5772
10.2478/aemnp-2019-0007
64a55ff4-0a71-491f-a4f4-9b2cd5a41431
1804-6487
4505468
027CE86F-9E75-44C3-A35E-E0C20BA4B693
Cleotomiroides ishikawachui
Yasunaga & Duwal
,
sp. nov.
(
Figs 33–35
,
74−76
,
192–195
)
Type material.
HOLOTYPE
: ♁,
INDONESIA
:
JAVA
:
Malang
,
Buring, S
07°59′40.0″–42.0″ E112°39′38.0″–39.1″,
513–518 m
alt.,
24 Aug 2005
(afternoon),
T. Ishikawa
(
TUAK
) (
AMNH
_
PBI 00380631
).
Differential diagnosis.
Recognized by its relatively large size; fuscous basic coloration contrasting with rusty apical half of the clavus (
Fig. 33
); conspicuous white fascia and maculae on hemelytron (
Figs 33, 35
); and form of male genitalia (
Figs 192−195
). This new species can be distinguished from two known congeners by the fuscous general coloration, broad hypophysis of the left paramere (
Fig. 193
), and elongate apical appendage and spinulate flap-like process of the endosoma (
Fig. 195
).
Description.
Male.
Body generally blackish brown, relatively small, myrmecomorphic; dorsal surface weakly shining, widely shagreened or roughened, with sparsely distributed, woolly semierect setae and with more sparsely distributed, longer, upright setae (
Fig. 35
). Head dull brown, oblique; eyes large; vertex narrowly carinate basally. Antenna dull yellowish brown, almost linear; segment II slightly incrassate toward apex; segments III and IV brown, weakly terete. Labium shiny dark brown, reaching subapical part of mesocoxa. Pronotum with a narrow collar, narrower than all antennal segments; pleura shiny fuscous, minutely rugose; ostiolar peritreme strongly protruding medially, with ivory white posterior margin of evaporative area. Hemelytron with a white, continuous fascia posterior to scutellum across clavus, corium and exocorium (
Fig. 33
); apical 1/4 of corium rusty; membrane smoky brown, except for pale, translucent base. Coxae and legs brown or darker; meso- and metacoxae widely pale brown; pretarsus with rather fleshy, apically convergent parempodia (
Fig. 76
). Abdomen wholly dark brown, somewhat shagreened (
Fig. 34
). Male genitalia (
Figs 192–195
): Genital segment with a ventral spine (
Fig. 192
). Left paramere with rather stout hypophysis (
Fig. 193
). Phallotheca almost straight, tapered, with a spine near apex (
Fig. 194
). Endosoma J-shaped, stout, with an elongate apical appendage and a weakly sclerotized, rounded flap-like process beside thick-rimmed secondary gonopore (
Fig. 196
).
Figs 40–55. Stridulatory device (MFP, except for FWS on 55) of Asian hallodapines observed by a compound microscope. 40–41 –
Hallodapus brunneus
(
Poppius, 1915
)
; 42–43 –
H. centrimaculatus
(Poppius, 1909)
; 44 –
H. fasciatus
(Poppius, 1909)
(Nakhon Nayok, Thailand); 45–46 –
H. kyushuensis
Miyamoto, 1965
; 47 –
H. spinosus
sp. nov.
; 48–50 –
H. ravenar
(Kirkaldy, 1902)
; 51 –
H. linnavuorii
Miyamoto, 1965
; 52 –
H. jingfui
sp. nov.
; 53 –
H. sibiricus
Poppius, 1912
(S. Primorsky, Russia); 54 –
Alloeomimella muiri
(
Schuh, 1984
)
; 55 –
Wygomiris paveli
sp. nov.
Measurements.
Male (
holotype
): Total length of body 3.33; head width including eyes 0.72; vertex width 0.21; lengths of antennal segments I–IV 0.30, 1.07, 0.60, 0.44; total labial length 1.28; basal width of pronotum 1.02; maximum width across hemelytron 1.05; and lengths of metafemur, tibia and tarsus 1.14, 1.70, 0.30.
Female.
Unknown.
Etymology.
Named after a Japanese heteropterist, Dr. Tadashi (nickname Chu)
Ishikawa
who collected the
holotype
specimen of this new species; a noun in genitive case.
Biology.
Unknown; Dr.
Ishikawa
collected the
holotype
female by sweep-netting an unidentified broadleaf tree.
Distribution.
Indonesia
:
Java
(this paper).