The Early Middle Eocene Wagon Bed Carpoflora Of Central Wyoming, U. S. A. Author Tiffney, Bruce H. Author Manchester, Steven R. text Fossil Imprint 2022 2022-08-26 78 1 51 79 http://dx.doi.org/10.37520/fi.2022.004 journal article 10.37520/fi.2022.004 2533-4069 7167719 Genus Alangium LAM. Text-fig. 6a–e M a t e r i a l. One chalcedony cast, DMNH EPI.47806. D e s c r i p t i o n. Bilocular endocarp represented by two locule casts arranged in the form of a sandwich ( Text-fig. 6c–e ) with the intervening septum also silicified. Endocarp wall missing. Locules elliptical in face view ( Text-fig. 6a, b ), 13.8–16.3 mm long, 11.5–12.9 mm wide, each 3–4 mm thick, one slightly larger than the other; fruit 9.3 mm thick perpendicular to the septum. Septum ca. 1.5 mm thick, with about 16 regularly spaced peripheral circular depressions corresponding to the central vasculature ( Text-fig. 6c–e ). D i s c u s s i o n. This distinctive cast was readily identified through comparison with similarly preserved specimens from the Clarno Formation of Oregon ( Manchester 1994 ). The Clarno specimens exhibit a range of preservation modes from permineralized examples with sufficient anatomical detail to verify the generic determination, to chalcedony locule cast “sandwiches” like the one figured here. The Wagon Bed specimen conforms to endocarps of section Marlea (ROXB.) BAILL. , which is also known from other Eocene localities including the Clarno Formation ( Manchester 1994 ), the London Clay ( Chandler 1961 ) and Messel ( Collinson et al. 2012 ) and from the Miocene Brandon Lignite ( Eyde et al. 1969 ). This specimen is distinctive in its large size in comparison to other Eocene species (length of 16 mm in this species vs. 10 mm in Clarno, 11 in London Clay, 6 mm in Messel). The modern genus includes about 20 species of small trees, shrubs and lianes of the Old World tropics, dominantly of southeast Asia ( Mabberley 2008 ). In sect. Marlea , A. scandens BLOEMB. is noted as a climber ( De Wilde and Duyfjes 2016 ).