Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales)
Author
Rio, Cédric Del
Author
Stull, Gregory W.
Author
Franceschi, Dario De
text
European Journal of Taxonomy
2020
2020-05-14
645
1
130
journal article
22093
10.5852/ejt.2020.645
1cd473d9-9d96-438f-bab4-29c5e56f82b8
3829651
Natsiatum herpeticum
Buch.
-Ham. ex Arn.
Fig. 19.18–19.26
Material examined
Specimen used for endocarp and fruit description
CHINA
• 1903;
H.B. Cale s.n.
;
P
[
MNHN-P-P04513511
].
Description
FRUIT. Elliptical to ovoid, apex slightly asymmetrical, black when mature. Epicarp strigose, with yellow simple hairs with granular ornamentation. Mesocarp
98–126 µm
thick when dry. Calyx persistent. Length 10.2–15.0 mm, width 7.6–10.0 mm, thickness
5.6 mm
.
ENDOCARP. Cream, elliptical to ovoid in lateral view, lenticular in transverse section, length ca
9 mm
, width ca
7 mm
, thickness ca
5.1 mm
. Keel surrounding the endocarp in the plane of symmetry, with a central channel. Apex asymmetrical and acute in lateral view; base rounded, symmetrical. Outer surface of the endocarp ridged, ridges faintly apparent, with at least two main longitudinal ridges; the median ridge reaching the point of the base. Ridges thin and rounded, with a median channel, delimiting a sparse reticulum with only 3–4 polygonal areoles visible. Vasculature resting in the channels of the ridges; endocarp primary vascular bundle semi-embedded within the endocarp wall in the channel of the keel. Endocarp wall
280–370 µm
thick excluding ridges (
432–531 µm
thick including ridges). Endocarp wall with two cell layers: outermost layer with 0–3 row(s) of isodiametric cells (mesocarp-endocarp cell transition), cells
15.1–19.7 µm
in diameter; innermost layer with 12–16 rows of periclinally oriented cells, cells
12.7–21.5 µm
in width. Locule surface unknown, lacunate.
Remarks
A figure from
Stull
et al.
(2011)
shows the endocarp of
Natsiatum herpeticum
Buch.
-Ham. ex Arn. better preserved with more areoles and freely ending ridgelets than seen here. We speculate that the development of the endocarp in our sample was halted, or the endocarp was decayed by fungal attacks (see Discussion for preservation issues).