An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion) Author Maddison, David R. text ZooKeys 2014 416 113 155 http://dx.doi.org/10.3897/zookeys.416.7706 journal article http://dx.doi.org/10.3897/zookeys.416.7706 1313-2970-416-113 CE0561FB5EE4498BA2C2EDF9B14F241D CE0561FB5EE4498BA2C2EDF9B14F241D Taxon classification Animalia Coleoptera Carabidae Subgenus Antiperyphus Jeannel, 1962 Antiperyphus Jeannel, 1962; type species Bembidium philippii Germain, by original designation. Remarks. As noted by Toledano (2008) , Jeannel's concept of Antiperyphus was polyphyletic, with at least Bembidion hirtipes , Bembidion ringueleti , Bembidion rufoplagiatum , and Bembidion uniforme Csiki belonging within Antiperyphanes . This is confirmed in part by my results (Fig. 8, Table 3). In addition, Bembidion engelhardti , Bembidion eburneonigrum , Bembidion tucumanum , and Bembidion germainianum are members of Nothonepha , not Antiperyphus . Of the species included in the subgenus by Jeannel (1962) , this leaves only the type species, Bembidium philippii (Fig. 2A). In addition, Bembidion peterseni Jensen-Haarup (1910) , from Mendoza, Argentina, can tentatively be placed here. I have examined a male syntype (in ZMUC), and it is similar in appearance to Bembidium philippii , although with much deeper and longer elytral striae. It is not a member of Nothonepha (as it lacks mesepisternal pits), nor is it a member of Antiperyphanes (it has a brush sclerite, and does not have the long flagellum characteristic of Antiperyphanes ). The internal sac of the male genitalia, although difficult to see because of the nature of the preparation, appears very similar to that of Bembidium philippii . B. philippii is common on sand shores of rivers in the provinces of Neuquen and Chubut in Argentina; it also occurs in Chile.