Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae) Author PRATHER, AYSHA L. text Zootaxa 2003 2003-08-29 275 1 1 214 https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1 journal article 5618 10.11646/zootaxa.275.1.1 9322f2b2-a384-40bf-9d57-d789c76ce5c4 1175­5334 5019235 89BC39FD-ABCC-4634-876D-8C1F4337C325 Phylloicus bicarinatus , new species Figs. 25 , 26 , 107 This species can be distinguished by the double keel or carinae of tergum X ( Fig. 25B ). The forewing pattern is similar to that of P. elegans and lituratus and maculatus , but in P. bicarinatus the setae forming the pattern are bright white, forming a crescent­shaped proximal band, in contrast to the ivory­yellow setae and the V­shaped proximal band of the other three species. Adult. Forewing length 9­9.9 mm, n = 28. Head chestnut brown. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax golden brown. Femora golden brown; foretibiae dark brown; mesotibiae dark brown; metatibiae dark brown; foretarsi chestnut brown; mesotarsi dark brown; metatarsi dark brown. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; black; with two transverse bands; proximal band white, extending from anterior to posterior wing margin; distal band white, beginning at anterior wing margin, at least 1/2 width of wing, widest and densest anteriorly; with single basal stripe, white ( Fig. 107 ). Hind wing basal brush present in male, dark brown. Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched, notch deep and round ( Fig. 25A, C ). Tergum IX without mesal ridge; posterior margin smoothly rounded; thinly sclerotized anteromesally ( Fig. 25B ); lateral ridge present; dorsal pleural setae approximately 8, ventral pleural setae absent ( Fig. 25A ). Preanal appendage less than 2/3 length of tergum X, narrowly elliptic, setae long, but not filamentous or longer than appendage ( Fig. 25A, B ). Tergum X without basal lobes; basodorsal process short and bifid; basolateral processes short, length less than or equal to diameter; apex, in lateral view, rounded, in dorsal view, notched, notch shallow; with paired mesal carinae ( Fig. 25A, B ). Harpago slightly tapered; peglike setae few, apical ( Fig. 25A, C ). Phallic endotheca with paired basolateral lobes, basolateral lobes tapered apically; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 25D, E ). Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII posterior margin entire; sternum VIII ( Fig. 26C ). Tergum IX with very short mesal ridge ( Fig. 26B ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 26A ). Tergum X appendage shorter than mesal lobe, base indistinct, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes long, at least twice diameter and often asymmetrical ( Fig. 26B ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 26A ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite truncate anteriorly, posterolateral projections absent; posterior sclerite ovoid ( Fig. 26A ). Holotype male: PERU : Madre de Dios : Manu Biosphere Res., Pakitza Bio. Sta., Quebrada Trompetero , trail 2, marker 15, 11°56'39”S , 71°16'59"W , 350 m , 3.vii.1993 , Blahnik & Pescador ( MHNJP ). Paratypes : BOLIVIA : La Paz : Río Alto Beni , Palos Blancos , 600 m , 11­15.i.1976 , Peña G. — 1 male ( NMNH ) ; Santa Cruz : Sara , 450 m , Steinbach — 1 male ( CMNH ) ; PERU : Cuzco : Quincemil , xi.1962 , Peña G. — 1 male ( CNC ) ; — 2 males ( NMNH ) ; Madre de Dios : Toma del Agua , Amazonia Lodge , 12°52'13”S , 71°22'34"W , 415 m , 29.vi.1993 , Blahnik & Pescador — 1 male ( NMNH ) ; — 1 male ( UMSP ) ; Manu Biosphere Res. , Pakitza Bio. Sta. , 11°56'00”S , 71°18'00"W , 350 m , 30.ix.1987 , Pogue — 1 male ( NMNH ) ; 01­13­03­99, 11°56'00”S , 71°18'00"W , 350 m , 2.x.1987 , Pogue — 1 male ( NMNH ) ; Aquajal , 11°56'00”S , 71°18'00"W , 250 m , 12.ix.1988 , Pogue — 2 males ( NMNH ) ; trail 1, marker 14 (1st stream), 11°56'00”S , 71°18'00"W , 250 m , 19­23.ix.1989 , Adams et al. — 1 male ( NMNH ) ; trail 1, marker 4 (near tents), 11°56'00”S , 71°18'00"W , 250 m , 8­22.ix.1989 , Adams et al. — 1 female , 3 males ( NMNH ) ; — 1 male ( MHNJP ) ; — 1 male ( UMSP ) ; trail 2, marker 18, 11°56'00”S , 71°18'00"W , 250 m , 12­23.ix.1989 , Adams et al. — 1 male ( MHNJP ) ; — 5 males ( NMNH ) ; Quebrada Paujil­Picoflor , trail 1, marker 13, 11°56'39”S , 71°16'59"W , 350 m , 2.vii.1993 , Blahnik & Pescador — 1 female ( NMNH ) ; 4­6.vii.1993 , Blahnik & Pescador — 1 female ( MHNJP ) ; — 1 female ( NMNH ) ; Quebrada Trompetero , trail 2, marker 15, 11°56'39”S , 71°16'59"W , 350 m , 3.vii.1993 , Blahnik & Pescador — 1 male ( MHNJP ) ; — 1 female , 2 males ( NMNH ) ; — 2 females ( UMSP ) . Distribution. Bolivia , Peru . Etymology. Bicarinatus , from the Latin bi­ , “two” and carinatus , “keeled,” referring to the double keel of tergum X.