Taxonomic review of the family Colatooeciidae Winston, 2005 (Bryozoa, Cheilostomata), with description of seven new species
Author
Almeida, Ana C. S.
Author
Souza, Facelucia B. C.
Author
Menegola, Carla M. S.
Author
Sanner, Joann
Author
Vieira, Leandro M.
carol.salmeida@gmail.com
text
Zootaxa
2014
2014-09-26
3868
1
1
61
journal article
4235
10.11646/zootaxa.3868.1.1
c0b3c91a-f100-4743-afcc-d4e77ff11d9e
1175-5326
4930657
ACC2AFCA-549B-4E2E-9E33-BF229FA3348A
Genus
Trematooecia
Osburn, 1940
Type
species.
Trematooecia aviculifera
(
Canu & Bassler, 1923
)
, now fixed under ICZN Article 70.3, misidentified as
Trematooecia turrita
(
Smitt, 1873
)
in the original designation by
Osburn (1940)
.
Diagnosis (revised).
Colony encrusting, uni- to multilaminar or erect and bilaminar. Autozooids heavily calcified. Frontal wall with frontal pseudopores and marginal areolar pores. Primary orifice transversely D-shaped or hoof-shaped. Secondary orifice formed by raised, rounded tubercles often obscuring the primary orifice. Suboral avicularium often present, seated within peristome near primary orifice. Interzooidal and adventitious avicularia with complete crossbar. Ovicell acleithral, with coarse surface calcification, ectooecium with a single membranous frontal area.
Remarks.
Trematooecia
was erected by
Osburn (1940)
to accommodate
L. turrita
Smitt, 1873
, designated
type
species, and
Trematooecia protecta
Osburn, 1940
. Re-examination of the specimens studied by
Osburn (1940)
from
Puerto Rico
(
Guanica
Harbor,
9–32 m
),
Tortugas Island
(
20–27 m
) and
Curaçao
(all USNM Acc. No. 208837), allow us to reassign these colonies as
T. aviculifera
, as suggested by
Banta & Carson (1977)
and
Winston (2005)
. The main difference between
T. aviculifera
and
C. turrita
is the ectooecium, with a circular uncalcified area in
T. aviculifera
and a longitudinal band of irregular pseudopores in
C. turrita
. Other differences pertain to the condyles (absent in
T. aviculifera
, present in
C. turrita
) and frontal avicularia (absent in
T. aviculifera
, dimorphic in
C. turrita
), but these characters seem to be variable within both
Trematooecia
and
Cigclisula
.
Owing to the misidentification of
T. aviculifera
as
C. turrita
, some other species were subsequently incorrectly attributed to
Trematooecia
, hence the genus has included species with two different
types
of ectooecium: (i) with an uncalcified central area, characteristic of
Trematooecia
, and (ii) with a median band of pseudopores, typical of
Cigclisula
. Thus, some species previously assigned to
Cigclisula
are here reassigned to
Trematooecia
, viz.
Trematooecia arborescens
(Canu & Bassler, 1928)
n. comb.
,
Trematooecia gemmea
(
Winston & Woollacott, 2009
)
n. comb.
,
Trematooecia hexagonalis
(Canu & Bassler, 1930)
and
Trematooecia verticalis
(
Maplestone, 1910
)
n. comb.