Amphibians and reptiles from the Neogene of Afghanistan
Author
Lapparent, France de
Author
Bailon, Salvador
Author
Augé, Marc Louis
Author
Rage, Jean-Claude
text
Geodiversitas
2020
2020-09-17
42
22
409
426
journal article
10.5252/geodiversitas2020v42a22
99b2432e-c149-4b82-a584-4b33d116d198
1638-9395
4447648
urn:lsid:zoobank.org:pub:07001ACA-EBDE-4256-BCB9-55E3159F81DC
? Gen. et sp. indet.
(
Figs 1A
;
2
AB)
LOCALITY AND AGE. — Sherullah 9, Khordkabul basin,
Afghanistan
, late Miocene, Late Vallesian-basal Turolian transition. MN10/11.
MATERIAL EXAMINED. —
One fragmentary humerus (AFG 1650), 1 incomplete vertebra (AFG 1651). This material was mentioned by
Roček & Rage (2000)
.
DESCRIPTION
The humerus is represented by a distal extremity which comprises the humeral ball, ulnar epicondyle and a short part of the diaphysis (
Figs 1A
;
2A
). The condyle is large and spherical. The cubital fossa is poorly limited laterally.The well-developed ulnar crest extends laterally; it is not deflected dorsally. It was likely short anteroposteriorly. The ulnar epicondyle is large whereas the radial epicondyle is damaged. A small radial crest slightly deflected ventrally is present. Because of the large size of the condyle and ulnar epicondyle, the distal extremity is widened transversely; moreover, it appears to be somewhat flattened dorsoventrally. Such a morphology recalls that of
Alytidae
of the
Discoglossinae
subfamily.
The vertebra is a presacral one (
Fig. 2B
). The anterior part of the centrum is damaged and covered by matrix. The vertebra is apparently opisthocoelous but this cannot be definitely confirmed. If it is really opisthoceoelous, then it probably belongs to the
Alytidae
. Its size is consistent with that of the humerus.
COMMENTS
The referral of these two specimens to the
Alytidae
is doubtful. It is based only on the overall morphology of an incomplete humerus and on a vertebra the main characteristic of which (opisthocoely) is not certain. The
Alytidae
is a primitive family whose earliest representative was recovered from the middle Jurassic of
England
(
Evans
et al.
1990
). Today they are known in Europe, west Asia, and northernmost Africa (
Frost
et al.
2006
). In the Cenozoic of Asia, apart from the possible alytid of Sherullah 9, the family was reported from the early Eocene of the Vastan Lignite Mine (
Bajpai & Kapur 2008
) and the middle Miocene lower Siwalik deposits (
Parmar & Jigmet 2014
), in
India
; from the Miocene of
Turkey
(
Claessens 1995
;
Sanchiz 1998
; Rückert-Ülkümen
et al.
2002) and in the Miocene of
Thailand
(
Rage & Ginsburg 1997
). However, this latter identification cannot be confirmed (
Roček & Rage 2000
).