A foundation monograph of Ipomoea (Convolvulaceae) in the New World Author Wood, John R. I. Author Munoz-Rodriguez, Pablo Author Williams, Bethany R. M. Author Scotland, Robert W. text PhytoKeys 2020 143 1 823 http://dx.doi.org/10.3897/phytokeys.143.32821 journal article http://dx.doi.org/10.3897/phytokeys.143.32821 1314-2003-143-1 F6F11A6EE4FF5A1885CEA2B60AE965A5 401. Ipomoea clavata (G. Don) Ooststr. ex J.F. Macbr. , Publ. Field Mus. Nat. Hist. Bot. Ser. 11 : 3. 1931. (Macbride 1931: 3) Calonyction clavatum G. Don , Gen. Hist. 4 : 264. 1838. (Don 1838: 264). Type. ECUADOR. Guayaquil, Ruiz & Pavon s.n. (holotype?BM ex Herb. Lambert, n.v.; isotypes MA, OXF). Convolvulus clavatus Pav. ex Choisy in A.P. de Candolle , Prodr. 9 : 346. 1845. (Choisy 1845: 346). Ipomoea lactescens Benth. , Pl. Hartweg. 120 . 1839. (Bentham 1839-57: 120). Type. ECUADOR. Guayaquil, K.T. Hartweg 676. (holotype K (not barcoded) ex Herb. Bentham, isotypes BM, K, OXF). Operculina hirsuta Standl., J. Washington Acad. Sci. 14(11): 242. 1924. (Standley 1924: 241). Type. EL SALVADOR. S. Calderon , 1338 (holotype US00111353, isotype NY). Ipomoea contrerasii L.O. Williams, Fieldiana , Bot. 32 (12): 189. 1970. (Williams 1970a: 189). Type. GUATEMALA. Peten , Arroyo Paxcaman, Uaxactun, E. Contreras 3640 (holotype LL00372563, isotypes F, K, MO, S). Type. Based on Calonyction clavatum G. Don Description. Twining perennial to c. 5 m; stems with long, white, stiff, spreading hairs. Leaves petiolate, 6-12 x 5-10 cm, ovate, sometimes shallowly 3-lobed or with a single lateral lobe, shortly acuminate and mucronate, cordate with rounded auricles, margin often undulate, glabrous, paler beneath, thin in texture, main veins prominent beneath; petioles 4-4.5 cm, pilose. Inflorescence of 1(-2)-flowered, axillary, pedunculate cymes; peduncles 0.4-2.5 cm; bracteoles 2 mm, lanceolate, caducous; pedicels 2-4 cm, darker than peduncle, conspicuously thickened upwards, glabrous; sepals subequal, 23-28 x 10 mm, broadly lanceolate, acuminate, glabrous, margin broad, scarious; corolla 7.5-11 cm long, glabrous, broadly funnel-shaped, the tube white, limb blue, deeply lobed. Capsules ovoid, c. 2 cm long, glabrous; seeds 10-13 x 5 mm, shortly tomentose but with long yellowish marginal hairs. Illustration. Figures 11F , 196 . Figure 196. Ipomoea clavata . A habit with flower B habit with buds C outer sepal D inner sepal E corolla opened out to show stamens F ovary and style G fruiting calyx H seed. Drawn by Rosemary Wise A from Hartweg 671; B from photo by Fuentes; C, D from Ruiz & Pavon s.n.; E-H from Fuentes & Miranda 10895. Distribution. Scattered in disturbed bushy places in areas of good rainfall at low altitudes up to just over 1000 m along the Andean chain from northern Bolivia to southern Mexico: BOLIVIA. La Paz : Sud Yungas, Alto Beni, R. Seidel 2455 (ARIZ, K, LPB); Madidi, A. Fuentes & T. Miranda 10895 (CTES, OXF, LPB, MO, USZ). PERU. Amazonas : Bagua, A. Gentry et al. 22840 (MO); ibid., F. de la Puente 2443 (CIP). Cajamarca : San Ignacio, J. Campos de la Cruz & O. Diaz 2373 (MO). Cusco : La Convencion , Maranura, L. Valenzuela et al. 3115 (MO, OXF). Junin : Satipo-La Merced, T. Croat & M. Sizemore 81993 (MO). Loreto : Via Nauta-Iquitos, C. Diaz & N. Jaramillo 1269 (MO). Pasco : Oxapampa, Pozuzo, R. Vasquez et al. 35838 (MO, OXF). San Martin : Rio Huallaga, Chazuta, G. Klug 4076 (BM, K, MO, S); ibid., Juan Jui, G. Klug 4308 (BM, K, MO, S). ECUADOR. Bolivar : La Chorrera, C. Jativa & C. Epling 017 (MO, S). El Oro : E. Asplund 15766 (S). Esmeraldas : J. Hudson 744 (MO, RB). Guayas : type of Ipomoea lactescens . Imbabura : Cotacachi, C.E. Ceron & C. Reyes 67397 (Q, QAP). Manabi : H. von Eggers 15458 (K). COLOMBIA. Cesar : Poponte, C. Allen 803 (MO). Cundinamarca : La Mesa-San Javier , Garcia Barriga 12048 (COL); Pacho, L. Uribe 1821 (COL). Valle : Hac. Hato Viejo, Vijes-Yotoco, J.E. Ramos 2752 (MO). COSTA RICA. Guanacaste, Santa Cruz, B. Hammel & I. Perez 24993 (CR, MO). NICARAGUA. Chontales, Rio San Juan, P. Shank & A. Molina 4595 (F, GH). EL SALVADOR. Ahuachapan , Area Protegida Santa Rita, J.M. Rosales 1951 (BM, MO); Santa Ana, San Diego-La Barra, D. Rodriguez et al. 2077 (BM). BELIZE. Corozal, P. Gentle 545 (F). GUATEMALA. Peten , Lago Peten Itza, B. Wallnoefer 9496 (K, MO, W); ibid., Laguna Macanche , R. Tun Ortiz 611 (F, MO). MEXICO. Campeche : Calamul, E. Martinez et al. 29709 (BM, MEXU); Tenabo, F. de la Puente 2939 (CIP). Guerrero : Montes de Oca, Vallecitos, G.B. Hinton 9676 (K, MO), Atoyac, Galeana, 10916 (K, MO), Mina , 11608 (K, MO); Juan R. Escudero, Tirra Colorada, H. Kruse 746 (IEB). Jalisco : La Huerta, Chamela, E. Lott & M. Butterwick 1518 (MO). Michoacan : El Camalote, J.C. Soto Nunez et al. 7126 (IEB, MEXU). Oaxaca : Asuncion Ixtaltepec, Cerro Timbon , A. Saynes & A. Sanchez 3416 (IEB). Quintana Roo : P. Moreno 536 (MEXU). Sinaloa : Concordia, A. Gonzalez s.n. [1/11/1994] (IEB). Veracruz : C.A. Purpus 7783 (GH). Yucatan : Izamal, G.F. Gaumer 984 (BM, E, F, K, MO). Note. An unmistakeable species because of its large blue flowers and pilose stems with very long white hairs.