Taxonomic notes on the genus Baetiella Uéno from China, with the descriptions of three new species (Ephemeroptera: Baetidae)
Author
Shi, Weifang
Author
Tong, Xiaoli
text
Zootaxa
2015
4012
3
553
569
journal article
10.11646/zootaxa.4012.3.9
6bce25b3-09f7-4a75-8687-08f3bd0ae131
1175-5326
253698
45CEB0B9-C7D2-4C1D-B399-3332D2F0CC5F
Baetiella bispinosa
(
Gose, 1980
)
(
Fig. 45
)
Pseudocloeon bispinosus
:
Gose, 1980
: 211
;
Baetiella bispinosa
:
Waltz & McCafferty, 1987
: 563
;
Tong & Dudgeon, 2000
: 143
Diagnosis.
Posterior margin of metanotum and abdominal terga I–II each with a single posteromedian protuberance, terga III–IX each with a pair of protuberances, the distance between protuberances gradually widened backwards, and distance between their bases on tergum III longer than the length of protuberance (
Fig. 45
).
Hind
wing pads vestigial. Basis of all legs with a single finger-like white coxal gill (cf.
Fig. 49
); claw with two rows of denticles, outer row with 8–9 acute denticles which successively increasing in length forwardly, inner row medially with 4–5 short and blunt denticles. Gills (or gill sockets) present on terga I–VII (
Fig. 45
). Median caudal filament reduced to one segment, cerci usually 1.5 times longer than body length.
Material examined.
CHINA
. Guangdong
: 16 nymphs, Yangchun, Bajia Town, Baishuihe River (
21.90°N
,
111.39°E
, alt.
110m
),
9.vi.2011
, leg. Weifang Shi; 1 nymph on slides, Lechang County, Jiufeng Town (
25.34°N
,
113.38°E
),
2.viii.2011
, leg. Huimin Zhong.
Distribution.
China
(Guangdong,
Hong Kong
);
Japan
.
Comments.
The specific name of this species first appeared in the diagnostic key to Japanese mayflies given by
Gose (1980)
who assigned it to the genus
Pseudocloeon
. However, Gose did not specifically state he was establishing a new species because he was neither designating nor describing the species and only provided a nymphal habitus illustration and some characters for the diagnostic key. According to the current Articles 10–16 and 71–76 of the International Code of Zoological Nomenclature (ICZN 1999), the name
Pseudocloeon bispinosus
Gose
should be regarded as an invalid name. Moreover, because
Gose (1980)
did not designate
type
specimens for this species, it should be also considered as
nomen dubium
. However, based on the second edition of the Code (ICZN 1962),
Pseudocloeon bispinosus
Gose
met the requirements for availability as a valid species because Gose’s paper was published before 1985.
Waltz & McCafferty (1987)
transferred it to the genus
Baetiella
.
Considering that the name
Baetiella bispinosa
(Gose)
has been traditionally accepted and widely used by mayfly workers, and until a
neotype
is established, it seems better to keep it as the current name, rather than to erect a
nomen novum
.