Three new species of penicillate millipedes from the Con Dao Islands of southeast Vietnam (Diplopoda, Lophoproctidae and Polyxenidae)
Author
Huynh, Cuong
text
Zootaxa
2020
2020-04-01
4759
1
1
30
journal article
10.11646/zootaxa.4759.1.1
f75b13f6-a898-42ff-93a4-2d177da2207f
1175-5326
3735855
8C3F94C9-3923-43F4-918A-6A1CA92F1F49
Genus
Monographis
Attems, 1907
Type
species:
Monographis kraepelini
Attems 1907
Monographis
Attems, 1907
;
Nguyen Duy-Jacquemin and Conde, 1967: 43
Other taxa included:
Within
Monographis
, twelve species have been described:
Monographis kraepelini
Attems, 1907
, from
Java
,
Indonesia
;
Monographis annandalei
(
Silvestri, 1948
)
and
Monographis mira
(
Turk
, 1947
)
from
India
;
Monographis demangei
Nguyen
Duy- Jacquemin and Condé, 1967, from
Vietnam
;
Monographis tamoyoensis
Condé and Nguyen Duy-Jacquemin, 1963
, from
Brazil
;
Monographis baihualingensis
Ishii and Yin, 2000
and
Monographis yunnanensis
Ishii and Yin, 2000
, from southern
China
;
Monographis queenslandica
Huynh and Veenstra, 2013
, from
Australia
;
Monographis dongnaiensis
Huynh and Veenstra, 2015
; and newly added
Monographis phuquocensis
Huynh and Veenstra, 2018c
, both were collected in
Vietnam
. Additionally, two subspecies,
Monographis kraepelini malayana
Nguyen Duy-Jacquemin and Condé, 1966
, and
Monographis kraepelini sabahna
Condé and Nguyen Duy-Jacquemin, 1984
, were collected from
Malaysia
.
Genus diagnosis
: Each eye 8 ommatidia. Body 10 tergites, 9 pleural projections. 13 pairs of legs. 8 antennomeres, VI longest, VIII shortest with 4 sensory cones. Antennomere VI with 3–5 thick bacilliform sensilla among 12–20 thin bacilliform sensilla, sensilla located in anterior position longer than posterior ones, arranged in simple crescent-shape. A row of backward-pointing minute setae located in the posterior margin of labrum. Anterior margin of labrum with acute triangular shape setae. Lateral palp of gnathochilarium 1.25 longer than medial palp. Gnathochilarium lateral palp with 11 conical sensilla. Spine on tarsus 1 and 2. Coxal glands on coxae 8th and 9th.
Genus remarks
: The genus
Monographis
is divided into two groups of species based on the arrangement of sensilla on antennomere VI in an either (1) simple crescent-shape with anterior sensilla longer than posterior ones (
M. kraepelini
,
M. annandalei
,
M. baihualingensis
,
M. demangei
,
M. mira
and
M. phuquocensis
) or (2) forming a triangular shape (
M. dongnaiensis
,
M. queenslandica
,
M. tamoyoensis
and
M. yunnanensis
).