On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae) Author Storozhenko, Sergey Yu. text Zootaxa 2021 2021-05-13 4970 1 106 118 journal article 6427 10.11646/zootaxa.4970.1.3 45d1d4f9-cd28-4759-bba9-4029f04f3f7e 1175-5326 4756017 9BF933DB-E764-42F8-94B6-257B91146E82 Genus Bettotania C. Willemse, 1933 Bettotania C. Willemse, 1933: 75 ( type species: Bettotania maculata C. Willemse, 1933 , by original designation); C. Willemse, 1956: 112 ; Ingrisch, 1989: 227 ; Otte, 1995: 277 ; Yin et al. , 1996: 96 . Paracelebesia Miller, 1935a: 697 ( type species: Paracelebesia festiva Miller, 1935 , by original designation); synonymized with Bettotania by C. Willemse (1956) . Description . Head in frontal view oval; frontal ridge narrow, almost reaching the clypeus. Face in lateral view reclinated; frontal ridge considerably projecting between the antennae and forming a widened rostrum ( Fig. 1 ). Ventral genicular lobes of the hind femora narrowly triangular with apex obtuse, without spine. Male 10th abdominal tergite with furculae. Male supra-anal plate long-oval, with a median basal impression, lateral margins with a pointed tooth somewhat behind the middle, apex rounded. Male cerci compressed, conical, apex pointed, inner side without tooth. Valves of ovipositor relatively long, with apex rounded and crenulate ( Fig. 1 ). Male genitalia: epiphallus divided and asymmetrical; bridge relatively broad; left lophus long, finger-shaped; right lophus small, rounded; ancorae absent; oval sclerites attached to epiphallus ( Fig. 19 ); apodemes broad; valves of cingulum short; basal valves of penis connected with apical ones by a sinuate, long, unbroken flexure ( Fig. 21 ). Species included . The genus consists of four species distributed in Malaysia (Peninsular Malaysia and Borneo Island).