New World species of Asymphyloptera Collin (Diptera: Empididae: Clinocerinae) Author Sinclair, Bradley J. text Zootaxa 2015 4048 4 553 564 journal article 10.11646/zootaxa.4048.4.6 12dd6a18-8481-4dc9-8727-8cfbfc138f7a 1175-5326 237775 FE00B390-0362-42F7-B6F8-67D35141E014 Asymphyloptera havasu sp. nov. ( Fig. 14 ) Type material . HOLOTYPE , ♂ labelled: “ USA : AZ:Grand Canyon NP/ CR#7, Elves Canyon [Chasm]/ 10– 21.v.1998 / J.F. MacDonald”; “ HOLOTYPE / Asymphyloptera / havasu / Sinclair [red label]” ( CNC ). PARATYPES : USA . Arizona : Same data as holotype ( 2 ♂ , 2 ♀, CNC ); Grand Canyon NP, CR#15, Fern Canyon, 10–21.v.1998 , J.F. MacDonald ( 1 ♂ , 2 ♀, CNC ); Grand Canyon NP, CR#3, Saddle Canyon, 10–21.v.1998 , J.F. MacDonald ( 1 ♂ , USNM ). Recognition . This mostly dark species may be distinguished by pale fore coxa (viewed anteriorly) and dark pleura; narrow, tapered surstylus with pointed apex; apex of phallus expanded, membranous and strongly bent anteriorly. Description . Male . Postpedicel globular with long, slender apical extension, arising subapically. Ocellar setae stout, similar in length to postpedicel extension. Labrum slightly shorter than clypeus; palpus brown, subequal to labrum, tapered to slender apex. Pleura dark brown with thin bluish pruinescence. Chaetotaxy very stout; 1 pprn short, very slender; 2 npl, lower slender and shorter; 4 dc, anterior dc offset; 2 apical sctl, slightly longer than prescutellar dc, lacking lateral sctl. Legs brown, fore coxae pale yellowish-brown (anterior view); fore and mid femora with antero- and posteroventral rows of pale, slender setae, subequal in length to width of femur; fore tibia with biserial row of erect setae. Wing length 1.7–2.0 mm; crossvein h distinct; base of wing with slender setulae along posterior margin. Halter pale, yellowish-brown. Male terminalia ( Fig. 14 ): Cercus with anterior lobe slightly produced; posterior lobe long, slender. Hypandrium prolonged as broad, paired postgonites; apical margin of postgonite not extended anteriorly, apex pointed. Epandrium narrow, with elongate setae on apical half. Surstylus shorter than cercus, slender, arched, tapered to pointed apex; not extending beyond phallus. Phallus straight; apex expanded, membranous, strongly bent anteriorly; apical margin hooked anteriorly; ejaculatory apodeme narrow, short, not expanded. Female . Similar to male except as follows: lacking long ventral setae on fore and mid femora and fore tibia. Terminalia not dissected. Etymology . Named after the Havasupai, a native American tribe that have lived for centuries in the Grand Canyon region; Havasu means “blue-green water” and pai “people”. Distribution . Known from several seepages along the Colorado River, collected by John MacDonald during a survey in Grand Canyon National Park ( Fig. 17 ). Remarks . This species was collected in deeply shaded madicolous habitats on the canyon walls, formed by cold streams flowing down narrow slot canyons ( Figs 6, 7 ). These sites are characterized by mosses and maidenhair ferns (MacDonald pers. comm. 2015).