Order Soricomorpha Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 220 311 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Sorex (Sorex) volnuchini Ognev 1922 Sorex (Sorex) volnuchini Ognev 1922 , Ann. Mus . Zool. Acad. St. Petersbourg, 22: 322 . Type Locality: Russia , Krasnodarskii kr., Adygeiskaya A.O. [middle course], r. Kisha (see Pavlinov and Rossolimo, 1987 ). Vernacular Names: Caucasian Pygmy Shrew . Subspecies: : Subspecies Sorex (Sorex) volnuchini subsp. volnuchini Ognev 1922 Subspecies Sorex (Sorex) volnuchini subsp. dahli Zagorodnyuk 1996 Distribution: S Russia and Caucasus States; Turkey and N Iran . Perhaps also Crimea , Ukraine . Conservation: IUCN – Lower Risk (lc). Discussion: Subgenus Sorex , S. minutus group. Formerly included in minutus but specimens from Caucasus have a slighly different karyotype (2n = 40, FN = 60) which led Kozlovsky (1973) and Sokolov and Tembotov (1989) to regard volnuchini as a full species. The karyotype of S. buchariensis is very similar ( Ivanitskaya et al., 1977 ). Zaitsev and Osipova (2003) were able to distinguish volnuchini morphologically from minutus ; they also documented Pleistocene records for volnuchini . Zagorodnyuk (1996 c ) described the population of Crimea as subspecies dahli , in contrast to the smaller minutus inhabiting mainland Ukraine . Kryštufek and Vohralík (2001) demonstrated that Turkish populations (except those from Thrace) represent S. volnuchini .