A New Genus of Mesomphaliine Tortoise Beetle (Coleoptera: Chrysomelidae), with Description of a New Flightless Species from Haiti Author Shin, Chulwoo text The Coleopterists Bulletin 2013 2013-12-31 67 4 521 531 http://dx.doi.org/10.1649/0010-065x-67.4.521 journal article 10.1649/0010-065x-67.4.521 1938-4394 10104027 Convexocoleus rileyi Shin , new species ( Figs. 1–6 , 14, 17, 20 ) Diagnosis. As for the genus (see above). Description. Male, length 9.9 mm, width 9.9 mm; female, length 11.0 mm, width 10.0 mm. Coloration mainly black; anterolateral and posterolateral Fig. 13. Consensus tree of 24 most parsimonious trees (Length = 150, CI = 0.42, RI = 0.69). Black circles indicate nonhomoplasious characters; white circles indicate homoplasies; character numbers are above black or white circles, character states are below the circles; taxa in boxes represent the flightless lineages. Only unambiguous characters are mapped. Figs. 14–22. Antennae: 14) Convexocoleus rileyi ; 15) Elytrogona quatuordecimmaculata ; 16) Botanochara impressa . Pretarsal claws, white outline shows the shape of the basal tooth on the pretarsal claw; arrow indicates the connected bases: 17) C. rileyi ; 18) E. quatuordecimmaculata ; 19) B. impressa . Anterior portion of prosternum, lateral view: 20) C. rileyi ; 21) E. quatuordecimmaculata ; 22) B. impressa . margins of pronotum orange to red; sub-basal, middle, and subapical regions of elytral explanate margins orange; sub-basal, postmedian, and subapical areas of disc also orange to red; legs ( Figs. 3, 6 , 17 ) black with pale brown setae; abdomen ( Figs. 3, 6 ) black with sublateral brown depressions on ventrites II–V. Antennomere ( Fig. 14 ) IV slender, slightly broader apically; antennomere V about 0.8 times as long as antennomere IV, broader apically; antennomeres VI–VIII each as long as antennomere V, broader than antennomere V; antennomeres IX–X quadrate, as long as antennomere V; antennomere XI slightly shorter than antennomere III. Pronotum ( Figs. 1, 4 ) hemispherical with disc and anterior margin slightly convex; lateral margin rounded, smoothly curved without angle at posterolateral margin; explanate margin broader posteriorly; basal line broadly rounded; disc distinctly delimited by black coloration and depression between disc and margin; discal surface smooth; lateral marginal area finely and sparsely punctate; ventral surface of anterior margin rugose ( Fig. 23 ). Prosternum ( Figs. 3, 6 , 20 ) with surface irregularly wrinkled; prosternal process slightly swollen, with carved mark on surface. Tarsomere I ( Figs. 1, 6 ) hemispherical; tarsomere II heart-shaped, 2 times as long as tarsomere I; tarsomere III bilobed, as long as broad, 2.5 times as long as tarsomere I; tarsomere IV as long as tarsomere III, 2.5 times as long as tarsomere I. Type Material. H o l o t y p e (m a l e), HAITI : Dept. Grand’ Anse: Jeremie , 0.5 km West of summit of Pic Macaya , 2,300 m , I-27-1985 , C. Fastie (deposited in TAMU , TAMU-ENTO x0996649). Paratype (female) , HAITI : Dept. Sud: Morne Formond N. slope, 1,950 m , I-5-1983 , C.A. Woods et al . (deposited in FSCA ) . Etymology. The specific epithet is a patronym honoring Mr. Edward G. Riley, entomology collection manager at Texas A&M University and specialist on the Chrysomelidae . He was the first to notice the distinctiveness of the unusual specimen, designated herein as holotype , from the TAMU collection.