Taxonomic and nomenclatorial revision of the Neotropical genus Phaeoxantha Chaudoir (Coleoptera: Cicindelidae) Author Moravec, Jiří 0000-0001-5294-6410 Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410 jirmor@quick.cz Author Dheurle, Charles 0000-0001-5294-6410 Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410 & 5 place Jenson, F- 52200 Langres, France. charles. dheurle @ wanadoo. fr; https: // orcid. org / 0000 - 0002 - 2580 - 6962 Corresponding author & Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410 jirmor@quick.cz text Zootaxa 2023 2023-12-14 5386 1 1 83 https://www.mapress.com/zt/article/download/zootaxa.5386.1.1/52481 journal article 10.11646/zootaxa.5386.1.1 1175-5334 10376586 9A5C0CC4-3D86-45BD-97FC-694A4E31A8B5 Phaeoxantha ( Euphaeoxantha ) wimmeri ( Mandl, 1958 ) . ( Figs 310–324 ) Megacephala ( Phaeoxantha ) wimmeri Mandl, 1958: 30 t, 30 d, pl. 3, figs. 6–7, 10. Phaeoxantha wimmeri : Guerra, Brzoska & Pearson 1997: 26 . Megacephala ( Phaeoxantha ) wimmeri : Wiesner 1992: 45 . Phaeoxantha wimmeri : Cassola & Pearson 2001: 11 . Phaeoxantha wimmeri : Wiesner 2020: 30 . Type locality. Bolivia : department of Santa Cruz , alt. 500 m . Type material . Holotype ( Fig. 313 ) and allotype in NHMB , holotype labelled: “ Bolivia oriental – / Santa Cruz 500m – / 10.10.54 Zischka” [bluish print/handwritten] // “ Megacephala / bucephala / W. Horn” [sic! handwritten] // “Meg. (Phaeox.) / testudinea [sic!] / det Dr. Ing. K. Mandl 1958 ” [handwritten/printed, additionally crossed] // “ Holotypus / Meg. (Phaecox.) / Wimmeri m. / det Dr. Ing. K. Mandl 1958 ” [red, handwritten/printed]. Other paratypes ( examined from photos Figs 310–311 ). 1 ♂ , 1 ♀ in SDEI with same first two labels as holotype [the second label crossed] // “ Paratypus Megac. (Phaeox.) / Wimmeri m / det Dr. Ing. K. Mandl 1958 ” [red, handwritten/printed]. 57 paratypes in ZSM (not examined). FIGURES 310–312. Phaeoxantha ( Euphaeoxantha ) wimmeri (Mandl) from Santa Cruz, Bolivia. 310–311—habitus: 310—♂; 24.4 mm, PT (SDEI); 311—♀, 24.3 mm, PT (SDEI); 312—labels of the female PT. Bars = 1 mm. Image credit Kevin Weissing. FIGURES 313–318. Phaeoxantha ( Euphaeoxantha ) wimmeri (Mandl) . 313—habitus, ♂, 25.8 mm, HT (NHMB), image credit Simona C. Ruffener (NHMB); 314—labels of the HT; 315–318—specimens from Yapacani in Santa Cruz, Bolivia (CCJM): 315–316—labrum: 315—♂; 316—♀; 317—head showing buccal appendages, ♂; 318—pronotum, ♀. Bars = 1 mm. FIGURES 319–324. Phaeoxantha ( Euphaeoxantha ) wimmeri (Mandl) , all from Yapacani in Santa Cruz, Bolivia (CCJM). 319–320—mandibles: 319—♂; 320—♀; 321—aedeagus; 322–324—elytra: 322–323—♂; 324—♀. Bars = 1 mm. FIGURE 325. Amazon Basin. Modified from a smaller map: https://upload.www.wikimedia.org/wikipedia/commons/0/02/ Amazonriverbasin_basemap.png). Other material examined . 2 ♀♀ in CCJM : “ Bolivia oriental. / Santa Cruz 500 m / 20.XI.1955 , Zischka lgt.” . 4 ♂♂ , 2 ♀♀ in COSJ , 4 ♂♂ , 2 ♀♀ in CCJM , 1 ♂ in CJVB : “ Bolivia : Santa Cruz depart. / 80 km NW of city Sta. Cruz / Yapacani env. 25.XI.2007 / O. Šafránek lgt.” . 2 ♂♂ , 4 ♀♀ in CJVB : ibid. except for: “ 21.I.2017 Kamil Klečka leg.” . 1 ♀ in CJVB : “ Bolivia : Santa Cruz / 80 km NW of city Sta. Cruz / El Torno env. / 21.I.2017 leg. Jakub Netušil . 1 ♂ , 2 ♀♀ in COSJ : “ Bolivia : Cochabamba depart. / cca 5 km N from Villa Tunari / S16°54´46´´ ; W65°24´24´´ 270 m / 22.XI.2012 , light trap / O. Šafránek et M. Amaya lgt.” . Differential diagnosis . Phaeoxantha ( Euphaeoxantha ) wimmeri is very similar to P. ( E .) klugii . As the elytral shape may be sometimes very similar in both these species due to the usual variability (but always shorter in P. ( E .) wimmeri ), and as also their aedeagi are very similar, one other distinguishing character is the shape of the right mandible in males of P. ( E .) wimmeri , which has the second tooth mostly tightly appressed to the terminal tooth and constantly markedly distant from the third tooth ( Figs 317 , 319 ). For other distinguishing characters see “Differential diagnosis” under P. ( E .) klugii above. Examined adult specimens are 22.5–26.1 mm long, 8.50–11.6 mm wide; the illustrated type specimens measure: length of the holotype (NHMB) 25.8 mm , width 11.3 mm ; paratypes (SDEI) length of male 24.4 mm , width 10.2; length of female 24.3 mm , width 9.70 mm . Distribution . Known only from Bolivia , hitherto only from the department of Santa Cruz ( Wiesner 2020 ), but three above-listed specimens from Villa Tunari come from the department of Cochabamba . Adults were taken on rather large sandy beaches of the rivers Yapacani ( Fig. 328 ) and Surutú flowing from the mountain range to more open areas; Tetracha ( Neotetracha ) fulgida ( Klug, 1834 ) and T. ( N .) thomsoniana (W. Horn, 1915 ) shared the same places (O. Šafránek, pers. com.). Pearson et al. (1999a) listed a great number of other records from the department of Santa Cruz , mostly from banks of Rio Surutú and Rio Piray (also spelled “Pirai”), from Yapacani as well as a number of specimens from Rio Ichilo on the border of the departments of Santa Cuz and Cochabamba .