Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities Author Brazidec, Manuel https://orcid.org/0000-0002-0860-8972 Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China manuel.brazidec@gmail.com Author Vilhelmsen, Lars https://orcid.org/0000-0002-5593-5722 Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark Author Boudinot, Brendon E. https://orcid.org/0000-0002-4588-0430 Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany Author Richter, Adrian https://orcid.org/0000-0001-5627-2302 Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan Author Hammel, Joerg U. https://orcid.org/0000-0002-6744-6811 Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany Author Perkovsky, Evgeny E. https://orcid.org/0000-0002-7959-4379 Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine Author Fan, Yong Fushun Amber Institute, Fushun 113005, China Author Wang, Zhen Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China Author Wu, Qiong Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China Author Wang, Bo State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China Author Perrichot, Vincent https://orcid.org/0000-0002-7973-0430 Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France text Arthropod Systematics & amp; Phylogeny 2024 2024-03-22 82 151 181 http://dx.doi.org/10.3897/asp.82.e111148 journal article http://dx.doi.org/10.3897/asp.82.e111148 1864-8312-82-151 43AC036E93CC4D79939A07DF54BE1A2D 8D187145BFCA55A1A73960CDE5BD124B 3.1.1.3.2. † Megacoxa janzeni sp. nov. Figures 2E and 7 Etymology. The specific epithet is a patronym honoring Jens-Wilhelm Janzen, former owner of the holotype specimen. Material studied. Holotype female IGR.BU-069; housed in the amber collection of the Geology Department and Museum of the University of Rennes , France (IGR). One paratype male NIGP203547; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences , Nanjing , China . Type locality. Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar. Age. Upper Albian to lower Cenomanian, mid-Cretaceous. Diagnosis. Median mesoscutal sulcus smooth (vs. crenulate in Megacoxa synchrotron gen. et sp. nov.) ; axillae not contiguous medially (vs. contiguous in † Megacoxa chandrahrasa gen. et sp. nov. and Megacoxa synchrotron gen. et sp. nov.) ; fore wing with very narrow marginal cell closed in straight line by Rs (vs. broad with Rs curved in † Megacoxa chandrahrasa gen. et sp. nov. ); trapezoid medial cell (Fig. 2E ; vs. narrow in † Megacoxa chandrahrasa gen. et sp. nov. ); posterior margin of metacoxa outcurved and bordered with large foveae (Fig. 7B ; vs. bordered with small foveae in † Megacoxa chandrahrasa gen. et sp. nov. and and straight posterior metacoxal margin in Megacoxa synchrotron gen. et sp. nov. ). Description. Body length for holotype 5.20 mm; for paratype 3.72 mm. - Head glabrous, globular, higher than long (length for holotype 0.81 mm; for paratype 0.69 mm; height for holotype 1.04 mm; for paratype 0.75 mm); compound eye oval, higher than long, ocular carina absent; vertex convex, shagreened; clypeus short; subantennal groove present; antenna short, less than half as long as body; scape as long as wide (length for holotype 0.19 mm); pedicel wider than long (length for holotype 0.12 mm); flagellomeres cylindrical, elongate; mandibles symmetrical, with three teeth; occipital carina minutely crenulate. - Mesosoma half as long as body length (length for holotype 2.28 mm; for paratype 1.62 mm); mesoscutum one third as long as mesosoma (length for holotype 0.92 mm; for paratype 0.63 mm; width for holotype 1.03 mm), shagreened, overhanging pronotum almost at right angle, divided by smooth median mesoscutal sulcus; parapsidal line present; axillae not contiguous medially, inner axillar groove crenulate; pronotum with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum carinate. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length for holotype 3.08 mm; for paratype 2.58 mm); R1, Sc+R and M+Cu pigmented, Rs present between r-rs and Rs+M, composing first submarginal cell, M and A pigmented, Rs closing in a straight line a very narrow medial cell, trapezoid medial cell composed by basal segments of M, Rs+M and Cu1 (shortest side) and by 1m-Cu, Cu pigmented to margin with a bend. - Legs with metacoxa enlarged, posteriorly curved, and notched just before trochanter insertion, bearing a row of large foveae along posterior margin; two mesotibial spurs present; metafemur swollen and bare; metatibia also swollen, with two spurs and bearing row of comb-like setae that continue on metabasitarsus; metatarsus almost as long as metatibia; metabasitarsus four times longer than following metatarsomeres. - Metasoma shorter than mesosoma, covered with short setae (length for holotype 2.12 mm; for paratype 1.41 mm); first tergite longest and second almost same length; third, fourth and fifth of similar length and shorter than first; last tergite twice as long as fifth; tergite and sternite inserted in bevel; hypopygium well-developed; ovipositor same length as metasoma (length for holotype 2.10 mm; OL/BL ratio 0.40), sheaths fully preserved in holotype, transversely striated.