Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood Author Frolov, Andrey V. Author Montreuil, Olivier Author Akhmetova, Lilia A. text Zootaxa 2016 4207 1 1 93 journal article 37549 10.11646/zootaxa.4207.1.1 cef35771-ff95-4a41-9af6-196dea899a64 1175-5326 200444 F2875582-31E2-496F-AEEF-1D657DD86C33 Madecorphnus niger Frolov, 2010 ( Figs. 15 A–F) Madecorphnus niger Frolov, 2010c : 1103 ; Frolov 2013a : 2 . Type material examined. Holotype ( Figs. 15 A–C), male, “Madagascar Est Perinet A. Peyrieras X-1972 / Museum Paris / Holotypus Madec. niger Frolov 2009 ” (MNHN). Paratype , male: “ Madagascar East Perinet A.Peyrieras XI-1973 ” ( MNHN ). FIGURE 14. Madecorphnus pauliani , holotype. Habitus in dorsal view (A), labels (B), aedeagus in lateral view and parameres in dorsal view (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F). Additional material examined. Male , “ MADAGASCAR : Province d'Antananarivo 3 km 41° NE Andranomay , 11.5 km 147° SSE Anjozorobe 1300m 5–13 Dec 2000 18°28′24″S , 47°57′36″E coll. Fisher , Griswold et al . California Acad of Sciences ex rotten log in montane rainforest BLF2537” ( CASC ) . Male , “ MADAGASCAR : Toamasina Parc National de Zahamena, Onibe River elev 780 m 21–23 February 2009 17°45′33″S , 048°51′17″E Calif. Acad. of Sciences coll. B.L.Fisher et al. yellow pan trap rainforest coll code: BLF22170” ( CASC ) . FIGURE 15. Madecorphnus niger . Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), internal sac of aedeagus (C—photograph, D—schematic representation of armature), distributional records (E), labels of holotype (F). Diagnosis. This species can be separated from other Madecorphnus species by the shape of the parameres and from many species also by darker color of the body. Description. Male. Body length 5.5–5.7 mm . Color black, mouthparts and legs brown ( Fig. 15 A). Right mandible slightly longer than left, without tooth basad of apex. Labrum trapezoidal, length about 1/4–1/ 5 width (in dorsal view). Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and a number of smaller setae. Frontoclypeus slightly depressed in the middle anteriorly, minutely punctate (punctures separated by more than 10 their diameters). Pronotum 1.8 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, minutely punctate. Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral callus. Elytra minutely punctate except for a few relatively large punctures in the middle of each elytron near stria 1. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur as long as 2 basal tarsomeres in mesothoracic legs and as long as 3 basal tarsomeres in metathoracic legs. Parameres curved and rounded in lateral view, without teeth but with concavities laterally ( Fig. 15 B). Endophallus armature consists of 3 similar highly-sclerotized, tooth-like sclerites ( Figs. 15 C, D). Female unknown. Variation. Except for body size, the examined specimens differ slightly in the length of the right mandible. Distribution and habitat. Madecorphnus niger is known from three localities in eastern Madagascar ( Fig. 15 E).