An overview on the genus Amiserica Nomura, 1974 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)
Author
Ahrens, Dirk
0000-0003-3524-7153
ahrens.dirk_col@gmx.de
Author
Liu, Wangang
0000-0003-3524-7153
ahrens.dirk_col@gmx.de
Author
Pham, Phu Van
0000-0001-9369-6649
phupham.iebr@gmail.com
Author
Fabrizi, Silvia
0000-0003-3524-7153
ahrens.dirk_col@gmx.de
text
Zootaxa
2021
2021-10-11
5050
1
1
63
http://dx.doi.org/10.11646/zootaxa.5050.1.1
journal article
3968
10.11646/zootaxa.5050.1.1
81282108-3550-4a28-83ca-15bb5b23881a
1175-5326
5560512
C0BF369D-F7CB-4139-A466-21E402A119B9
Amiserica rufidula
(Nomura)
group
Checklist (up to this paper):
Amiserica rufidula
Nomura, 1974: 84
.
Amiserica babai
Kobayashi, 1988
b: 53
. (uncertain assignment)
Amiserica shizumui
Kobayashi, 1980: 17
. (uncertain assignment)
Remarks.
After the revision of extensive material of the genus
Maladera
Mulsant & Rey, 1871
from
China
(
Fabrizi
et al.
2021
), we could recognize
Amiserica loi
Kobayashi, 1991
(based on the original description and without the need of examination of the
type
material) as representative of the subgenus
Eumaladera
Nomura, 1967
:
Maladera
(
Eumaladera
)
loi
(
Kobayashi, 1991
)
,
new combination
. This species is excluded from
Amiserica
based on its characteristic genital morphology (
Kobayashi 1991
:
Fig. 7a–c
).
The species group assignment of the other included species is rather complicated.
Amiserica shizumui
Kobayashi, 1980
(see
Ahrens 2002
) and
Amiserica babai
Kobayashi, 1988
lack the membraneous sacs of parameres. However, in the first species, the metatibia also possesses a long serrated line beside the dorsal margin and a serrated line close to the anterior margin of metafemur while the anterior margin is robustly margined. Possibly both species represent isolated lineages. We had no material available for
Amiserica babai
.
Key to the species of the
Amiserica rufidula
(Nomura)
group (
♂♂
)
1 Antennal club composed of 3 antennomeres................................................................. 2
- Antennal club composed of 4 antennomeres................................................................ 4
2 Metatibia lacking serrated line.......................................................................... 3
- Metatibia lacking serrated line present. Left dorsolateral apical process of phallobase..........
A. rufidula
Nomura, 1974
3 Dorsolateral processes of phallobase subsymmetrical.........
A. zhongtiaoshanensis
Ahrens, Fabrizi & Liu
,
new species
- Dorsolateral processes of phallobase distinctly asymmetrical........
A. sacculiforceps
Ahrens, Fabrizi & Liu
,
new species
4 Apex of phallobase with at least short dorsolateral processes, phallobase mesoventrally enlarged...................... 9
- Apex of phallobase without long dorsolateral processes, phallobase mesoventrally not enlarged....................... 5
5 Antennal club long, twice as long as remining antennomeres combined. (
Vietnam
).........................................................................................
A. basisymmetrica
Ahrens, Fabrizi & Liu
,
new species
- Antennal club very long, three times as long as remaining antennomeres combined.................................. 6
6 Metatibia widest at middle and moderately wide, basal group of spines reduced to a single robust seta.................. 8
- Metatibia widest at apex and narrow, with a distinct basal group of spines at middle. Basal lobe of right paramere shorter and convexly rounded at mesal apex.......................................................................... 7
7 Metatibia with serrated line beside dorsal margin. Parameres at apex weakly curved dorsally, lateral lobe of left paramere attached to the paramere (
Taiwan)
..................................................
A. antennalis
(
Nomura, 1974
)
- Metatibia without serrated line beside dorsal margin. Parameres at apex strongly curved dorsally, lateral lobe of left paramere distinctly divergent from the paramere (
Yunnan
)...............
A. pseudoantennalis
Ahrens, Fabrizi & Liu
,
new species
8 Apex of phallobase entirely without dorsolateral processes. Basal lobe of right paramere long and sharply pointed at mesal apex. (
Vietnam
).................................................
A. strnadi
Ahrens, Fabrizi & Liu
,
new species
- Apex of phallobase with a short lateral process on the right side. Basal lobe of right paramere short and convex at apex. (
Vietnam
)...................................................
A. eucurviforceps
Ahrens, Fabrizi & Liu
,
new species
9 Metatibia with serrated line beside dorsal margin. Dorsolateral phallobasal processes shorter than half length of parameres. 10
- Metatibia without serrated line beside dorsal margin. Dorsolateral phallobasal processes more than half length of parameres. .................................................................................................. 11
10 Mesoventral extension of phallobase in lateral view of equal height. Right paramere short and weakly bent. Right dorsolateral processes short, not exceeding the base of right paramere..........
A. diaocangshana
Ahrens, Fabrizi & Liu
,
new species
- Mesoventral extension of phallobase in lateral view highest at middle. Right paramere longer and strongly bent. Right dorsolateral processes longer, distinctly exceeding distally the base of right paramere..............................................................................................
A. longxinensis
Ahrens, Fabrizi & Liu
,
new species
11 Narrowed portion of left phallobasal dorsolateral apical process straight and about half as long as the process itself....... 12
- Narrowed portion of left phallobasal dorsolateral apical process short and curved, much less than half as long as the process itself.............................................................................................. 13
12 Right paramere strongly narrowed at middle (lateral view). Right dorsolateral apical process of phallobase wide.........................................................................
A. incisa
Ahrens, Fabrizi & Liu
,
new species
- Right paramere even in width (lateral view). Right dorsolateral apical process of phallobase narrow..............................................................................
A. pseudoincisa
Ahrens, Fabrizi & Liu
,
new species
13 Left paramere subparallel (in lateral view)................................................................ 14
- Left paramere convexly narrowed from basal third towards apex and base (in lateral view)....................................................................................
A. funiushanica
Ahrens, Fabrizi & Liu
,
new species
14 Dorsolateral apical process of phallobase covering less than half of parameres. Right paramere slightly bent dorsally at middle (lateral view)........................................................................................ 15
- Dorsolateral apical process of phallobase covering more than half of parameres. Right paramere straight (lateral view)....................................................................
A. piaoac
Ahrens, Fabrizi & Liu
,
new species
15 Left paramere almost straight (lateral view). The right dorsolateral apical process of phallobase is more robust and more rounded at apex............................................
A. fengyangensis
Ahrens, Fabrizi & Liu
,
new species
- Left paramere curved upward (lateral view). The right process of the dorsolateral apical processes of phallobase is shorter and narrower than the left one.......................................
A. jiuhuensis
Ahrens, Fabrizi & Liu
,
new species