<document id="7FDFDB2264A9D47ECE064ED919AE6482" ID-ISSN="0024-4082" ID-ZooBank="7A6A530-3EBA-4AF2-94E2-E9B8A1D07459" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738713251128" checkinUser="plazi" docAuthor="Roggero, Angela, Dierkens, Michael, Barbero, Enrico &amp; Palestrini, Claudia" docDate="2017" docId="03DC6B6BFFF1FFD7FF07FDDDFEB8FB60" docLanguage="en" docName="zoj.12498.pdf" docOrigin="Zoological Journal of the Linnean Society 180" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Hamonthophagus fallax" docType="treatment" docVersion="2" lastPageNumber="314" masterDocId="FFE51313FFE1FFC7FFA1FFD5FF82FFF0" masterDocTitle="Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)" masterLastPageNumber="320" masterPageNumber="298" pageNumber="314" updateTime="1738713669289" updateUser="ExternalLinkService">
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<mods:title id="384AA72E8870870BB62439D9A87E536D">Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)</mods:title>
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<mods:namePart id="369974C3BEF9BA992F6AA4CB148D0F97">Roggero, Angela</mods:namePart>
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<mods:namePart id="8BDEA578CA985DFDB0BDDE1D1A0383E7">Dierkens, Michael</mods:namePart>
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<mods:namePart id="8DE375DA795828D8C7CF6577009848AC">Palestrini, Claudia</mods:namePart>
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<mods:title id="24683BA8944EB9AFE375989EA176BCCC">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="88C12E0FFC7B09040EB4EF5A6CD6ADFF">2017</mods:date>
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<treatment id="03DC6B6BFFF1FFD7FF07FDDDFEB8FB60" ID-DOI="http://doi.org/10.5281/zenodo.14807218" ID-Zenodo-Dep="14807218" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF1FFD7FF07FDDDFEB8FB60" httpUri="http://treatment.plazi.org/id/03DC6B6BFFF1FFD7FF07FDDDFEB8FB60" lastPageNumber="314" pageId="16" pageNumber="314">
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<paragraph id="8BCADA7DFFF1FFD7FF07FDDDFC86FDED" blockId="16.[166,772,515,573]" box="[166,772,515,542]" pageId="16" pageNumber="314">
<heading id="D0826D11FFF1FFD7FF07FDDDFC86FDED" box="[166,772,515,542]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="314" reason="2">
<taxonomicName id="4C75A1FEFFF1FFD7FF07FDDDFC86FDED" ID-CoL="3JJ8V" authority="(D' ORBIGNY, 1913: 471)" baseAuthorityName="D' ORBIGNY" baseAuthorityPageNumber="471" baseAuthorityYear="1913" box="[166,772,515,542]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B901066FFFF1FFD7FF07FDDDFE5FFDEB" box="[166,477,520,541]" italics="true" pageId="16" pageNumber="314">
<smallCapsWord id="8D2C4CA1FFF1FFD7FF07FDDDFE00FDEB" baselines="536,535" box="[166,386,520,541]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hamonthophagus" pageId="16" pageNumber="314">HAMONTHOPHAGUS</smallCapsWord>
<smallCapsWord id="8D2C4CA1FFF1FFD7FE2AFDDEFE5FFDEB" baselines="535" box="[395,477,523,539]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="fallax" pageId="16" pageNumber="314">FALLAX</smallCapsWord>
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(
<smallCapsWord id="8D2C4CA1FFF1FFD7FE51FDDEFD87FDED" baselines="535" box="[496,517,515,541]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="d" pageId="16" pageNumber="314">D’</smallCapsWord>
<smallCapsWord id="8D2C4CA1FFF1FFD7FDA4FDD0FDE9FDEC" baselines="535,536" box="[517,619,517,541]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Orbigny" pageId="16" pageNumber="314">ORBIGNY</smallCapsWord>
, 1913: 471)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36F89F6FFF1FFD7FECAFDF7FEB8FB60" pageId="16" pageNumber="314" type="description">
<paragraph id="8BCADA7DFFF1FFD7FECAFDF7FDBDFDCC" blockId="16.[166,772,515,573]" box="[363,575,546,573]" pageId="16" pageNumber="314">
(
<figureCitation id="134EC6F8FFF1FFD7FED5FDF1FE47FDCC" box="[372,453,546,572]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="16" pageNumber="314">
<smallCapsWord id="8D2C4CA1FFF1FFD7FED5FDF1FE2BFDCB" baselines="566,567" box="[372,425,548,572]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="16" pageNumber="314">FIGS</smallCapsWord>
2
</figureCitation>
, 
<figureCitation id="134EC6F8FFF1FFD7FE75FDF6FE64FDCD" box="[468,486,547,573]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="16" pageNumber="314">7</figureCitation>
, 
<figureCitation id="134EC6F8FFF1FFD7FE54FDF7FD84FDCC" box="[501,518,546,572]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="16" pageNumber="314">8</figureCitation>
, 
<figureCitation id="134EC6F8FFF1FFD7FDB7FDF7FDB7FDCC" box="[534,565,546,572]" captionStart="Figure 10" captionStartId="9.[148,227,1743,1762]" captionTargetBox="[373,1197,1135,1709]" captionTargetId="figure-119@9.[373,1197,1135,1709]" captionTargetPageId="9" captionText="Figure 10. Vagina and receptaculum seminis of the species of the genus Hamonthophagus (AC, BI, DE, FA and LA). Scale bar = 0.5 mm." pageId="16" pageNumber="314">10</figureCitation>
)
</paragraph>
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<emphasis id="B901066FFFF1FFD7FF03FDA6FED1FD77" box="[162,339,626,648]" italics="true" pageId="16" pageNumber="314">Type material:</emphasis>
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<typeStatus id="54CE64DFFFF1FFD7FEC3FDA6FE52FD78" box="[354,464,627,648]" pageId="16" pageNumber="314" type="lectotype">Lectotype</typeStatus>
here designated: (male) 
<collectingCountry id="F3629AEDFFF1FFD7FF03FD44FE91FD56" box="[162,275,657,678]" name="Malawi" pageId="16" pageNumber="314">MALAWI</collectingCountry>
: [Nyassa 
<collectingRegion id="49B1149FFFF1FFD7FE3DFD44FE6AFD57" box="[412,488,657,679]" country="Malawi" name="Zomba" pageId="16" pageNumber="314">Zomba</collectingRegion>
haut Chire=] 
<taxonomicName id="4C75A1FEFFF1FFD7FD14FD44FF5DFD35" authority=", Shire" authorityName="Shire" class="Insecta" family="Scarabaeidae" genus="Zomba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus">Zomba, Shire</taxonomicName>
river upper course, 
<location id="8EAA8CA6FFF1FFD7FE4CFD7AFD0EFD35" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF1FFD7FF07FDDDFEB8FB60:8EAA8CA6FFF1FFD7FE4CFD7AFD0EFD35" box="[493,652,687,709]" country="Malawi" name="Malawi Lake" pageId="16" pageNumber="314" stateProvince="Zomba">Malawi Lake</location>
[
<collectionCode id="ED6442B8FFF1FFD7FD02FD65FD7CFD35" box="[675,766,688,709]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="314">MNHN</collectionCode>
]
</materialsCitation>
. 
<materialsCitation id="3B1DD020FFF1FFD7FF03FD1BFEA1FCB0" collectionCode="MNHN" country="Tanzania" location="Dar-es-Salaam" pageId="16" pageNumber="314" specimenCount="1" stateProvince="Dar es Salaam" typeStatus="paralectotype">
<typeStatus id="54CE64DFFFF1FFD7FF03FD1BFEC0FD14" box="[162,322,718,740]" pageId="16" pageNumber="314" type="paralectotype">Paralectotype</typeStatus>
: (female) 
<collectingCountry id="F3629AEDFFF1FFD7FE6DFD1BFDD0FD13" box="[460,594,718,739]" name="Tanzania" pageId="16" pageNumber="314">TANZANIA</collectingCountry>
[=Afrique Or Alem]: 
<collectingRegion id="49B1149FFFF1FFD7FEBAFD39FE46FCF2" box="[283,452,748,770]" country="Tanzania" name="Dar es Salaam" pageId="16" pageNumber="314">Dar-es-Salaam</collectingRegion>
[
<collectionCode id="ED6442B8FFF1FFD7FE58FD38FDD6FCF2" box="[505,596,749,770]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="314">MNHN</collectionCode>
]. For the morphological account, please refer to the original description
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.
</paragraph>
<paragraph id="8BCADA7DFFF1FFD7FF03FCB2FEB8FB60" blockId="16.[162,777,871,1168]" pageId="16" pageNumber="314">
<emphasis id="B901066FFFF1FFD7FF03FCB2FDBEFC8C" box="[162,572,871,892]" italics="true" pageId="16" pageNumber="314">
Geographical distribution (
<figureCitation id="134EC6F8FFF1FFD7FE7CFCB2FDACFC8C" box="[477,558,871,892]" captionStart="Figure 14" captionStartId="12.[162,241,1389,1408]" captionTargetBox="[391,1208,203,1361]" captionTargetId="figure-188@12.[388,1212,198,1356]" captionTargetPageId="12" captionText="Figure 14. Distribution of the species (see Appendix 3, Supporting Information for a list of localities) with the Olson et al. (2001) terrestrial biomes classification." pageId="16" pageNumber="314">Fig. 14</figureCitation>
):
</emphasis>
The species has been described from 
<collectingCountry id="F3629AEDFFF1FFD7FE3EFC50FE76FC6B" box="[415,500,901,923]" name="Malawi" pageId="16" pageNumber="314">Malawi</collectingCountry>
and 
<collectingCountry id="F3629AEDFFF1FFD7FD9BFC50FD25FC6B" box="[570,679,901,923]" name="Tanzania" pageId="16" pageNumber="314">Tanzania</collectingCountry>
, and at present is recorded from 
<collectingCountry id="F3629AEDFFF1FFD7FDAEFC71FD00FC49" box="[527,642,932,953]" name="Botswana" pageId="16" pageNumber="314">Botswana</collectingCountry>
, 
<collectingCountry id="F3629AEDFFF1FFD7FD00FC71FC81FC4A" box="[673,771,932,954]" name="Burundi" pageId="16" pageNumber="314">Burundi</collectingCountry>
, 
<collectingCountry id="F3629AEDFFF1FFD7FF03FC17FDA8FC28" box="[162,554,962,984]" name="Democratic Republic of the Congo" pageId="16" pageNumber="314">Democratic Republic of Congo</collectingCountry>
, 
<collectingCountry id="F3629AEDFFF1FFD7FDE4FC16FD13FC28" box="[581,657,963,984]" name="Kenya" pageId="16" pageNumber="314">Kenya</collectingCountry>
, 
<collectingCountry id="F3629AEDFFF1FFD7FD0DFC17FC86FC28" box="[684,772,962,984]" name="Malawi" pageId="16" pageNumber="314">Malawi</collectingCountry>
, 
<collectingCountry id="F3629AEDFFF1FFD7FF03FC34FE85FC07" box="[162,263,993,1015]" name="Namibia" pageId="16" pageNumber="314">Namibia</collectingCountry>
, 
<collectingCountry id="F3629AEDFFF1FFD7FEBAFC34FE04FC07" box="[283,390,993,1015]" name="Tanzania" pageId="16" pageNumber="314">Tanzania</collectingCountry>
and 
<collectingCountry id="F3629AEDFFF1FFD7FE70FC34FDA8FC07" box="[465,554,993,1015]" name="Zambia" pageId="16" pageNumber="314">Zambia</collectingCountry>
. The record from Graaf-Reinet (
<collectingRegion id="49B1149FFFF1FFD7FEEBFBD5FDD9FBE6" box="[330,603,1024,1046]" country="South Africa" name="Eastern Cape" pageId="16" pageNumber="314">Eastern Cape province</collectingRegion>
, 
<collectingCountry id="F3629AEDFFF1FFD7FDCAFBD5FC83FBE6" box="[619,769,1024,1046]" name="South Africa" pageId="16" pageNumber="314">South Africa</collectingCountry>
) is very interesting, but needs to be confirmed by further records. A detailed list of the localities is given in the Appendix 3 (Supporting Information).
</paragraph>
</subSubSection>
</treatment>
</document>