Synopsis of Miridae (Hemiptera: Heteroptera) in Atlantic Forest Dominion, Espírito Santo State, Brazil: keys, diagnoses, new species, plant associations, and geographic distribution. Part I: Bryocorinae, Cylapinae and Deraeocorinae. Author Ferreira, Paulo Sérgio Fiuza Universidade Federal de Viçosa, Departamento de Entomologia (Full Professor retired), Avenida Peter Henry Rolfs, s / n, Viçosa, MG 36570 - 900, Brazil. Author Martins, David Dos Santos Instituto Capixaba de Pesquisa, Assistência Técnica e Extensão Rural, Rua Afonso Sarlo, 160, Bento Ferreira, Vitória / ES 29052 - 010, Brazil. Author Ferreira, Luciano Santana Fiuza Universidade Federal de Viçosa, Bacharelando em Cooperativismo, Departamento de Economia Rural, Edifício Edson Potsch Magalhães-R. Purdue, Campus Universitário, Viçosa, Minas Gerais, Brazil. Author Fornazier, Maurício José Instituto Capixaba de Pesquisa, Assistência Técnica e Extensão Rural (Incaper), Domingos Martins, ES, Brazil. text Zootaxa 2024 2024-06-12 5468 2 201 254 http://dx.doi.org/10.11646/zootaxa.5468.2.1 journal article 298417 10.11646/zootaxa.5468.2.1 ad0e6247-faa5-4276-a51e-0834afe872cf 1175-5326 11615431 4AD95CF2-297D-40F4-9DA4-71EC980499AA Pycnoderes quadrimaculatus Guérin-Menèville ( Fig. 6-P ) Pycnoderes quadrimaculatus Guérin-Menèville, 1857:404 ; Schuh 2003–2013 (online catalog, hosts, distribution); Knight 1968:77 (biology); Capriles 1969:20 (description); Carvalho & Ferreira 1972:162 (list); Alayo 1974:21 (diagnosis); Carvalho 1980B:647 (diagnosis, type specimen); Hernández & Henry 2010:41 (synonymy, diagnosis, color habitus, male genitalia, host, information); Alvarez-Zapata 2022:134 (diagnosis, key to species of Colombia , host, distribution). Diagnosis. Overall coloration is black, with areas of pale white and yellowish brown. Body oval or oblong, with silvery spots on the hemelytra. The head with vertex; front with a median line; and the clypeus with black apex. Pronotum disc greatly inflated with weakly grooved on median area. Hemelytra with an enlarged embolium and a pale subbasal spot about twice the length of the smaller apical spot. Male genitalia as figure. 11-K. Plant Associations. Alvarez-Zapata et al. (2022): Amaranthus sp. ( Amaranthaceae ) ( Ferreira et al. 2001 A), Zantedeschia aethiopica (L.) Spreng. ( Araceae ) (Nogueira et al. 2019A), Lactuca sativa L. ( Asteraceae ), Brassica rapa L. ( Brassicaceae ) (USDA 1941), Ipomoea sp. ( Convolvulaceae ) ( Hernández & Henry 2010 ), Citrullus lanatus (Thunb.) Matsum. & Nakai ( Cucurbitaceae ) ( Wehrle 1935 ), Cucumis dipsaceus Ehrenb. Ex Spach ( Cucurbitaceae ) ( Zimmerman 1948 ), Cucumis melo L. ( Cucurbitaceae ) ( Wehrle 1935 ), Cucurbita maxima Duchesne ( Cucurbitaceae ) ( Ferreira et al. 2001 ), Cucurbita pepo L. (Cucurbitacae), Cucurbitasativus ” ( Cucurbitaceae ) ( Hernández & Henry 2010 ), Phaseolus vulgaris L. ( Fabaceae ) ( Ferreira et al. 2001 ), Vigna unguiculata (L.) Walp. ( Fabaceae ) (Nogueira et al. 2019A), Carya illinoinensis (Wangenh.) K. Koch ( Juglandaceae ) (Life 2019), Saccharum spp . ( Poaceae ) ( Salazar et al. 2017 ) and Portulaca sp. ( Portulacaceae ) ( Zimmerman 1948 ). Distribution. Alvarez-Zapata at al. (2022): Argentina ( Carpintero & Carvalho 1993 ), Brazil : Espírito Santo (Jerônimo Monteiro ( Vélez et al. 2020 ), Domingos Martins, Aracruz, Mato Grosso , Minas Gerais , Piaui ) Colombia ( Nariño , Putumayo ) ( Carvalho & Afonso 1977 ; Alvarez-Zapata at al. 2022), Costa Rica ( Hemp & Konrad 2000 ), Cuba ( Guérin-Menèville 1857 ) , Dominican Republic (Carvalho 1951A), Guatemala ( Carvalho & Afonso 1977 ), Honduras (Carvalho 1951A), Mexico (Alvarez-Zapata at al. 2022), Nicaragua , Panama ( Carvalho & Afonso 1977 ), Peru ( Carvalho 1991 ; Carvalho & Ferreira 1972 ), Puerto Rico , Surinam (Carvalho & Rosas 1965), U.S.A. ( Uhler 1894 ; Zimmerman 1948 ), Venezuela ( Cazorla-Perfetti & Morales-Moreno 2019 ), West Indies (Van Duzze 1917; Uhler 1894 ).