Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia Author Cárdenas-Bautista, Johann Stephens Author Parada-Alfonso, Jenny Andrea Author Carvajal‑Cogollo, Juan E. text Check List 2020 2020-07-03 16 4 821 839 http://dx.doi.org/10.15560/16.4.821 journal article 10.15560/16.4.821 1809-127X Dichotomius quinquelobatus ( Felsche, 1901 ) Materials examined. COLOMBIA • 2♀ , 21.40 mm (mean); Meta , Villavicencio , Vereda Buenavista , “El Porvenir” Farm ; 04°08′32.33″N , 073°40ʹ48.44″W ; 1095 m a.s.l. ; 23 Dec. 2018 ; Cárdenas Johann and Parada Jenny leg.; high secondary forest; Bvbt 7; UPTC-In-00061 . • 1♂ , 21.40 mm (mean); Meta , Villavicencio , Vereda El Carmen , Caño Blanco–Caño Buque ; 04°08ʹ21.08″N , 073°40ʹ08.76″W ; 761 m a.s.l. ; 30 Apr. 2019 ; Cárdenas Johann and Parada Jenny leg.; riparian forest; Crbrt 5; UPTC-In-00062 . Identification. This species can be distinguished from other Dichotomius species , in the case of males, by the head with transverse roughness of the clypeus and granular punctation in the frons and gena; clypeus with a transversely flattened horn; pronotum with a strong flat slope with marked punctation and roughness with five protuberances in the form of small horns located along the posterior margin of the slope, the central horn and the lateral ends are slightly larger than those near the central horn. In females, the head has a conical protuberance towards the vertex and frons region; pronotum with a pronounced declination towards the anterior medial region with two previously projected medial tubercles towards the apex and two smaller lateral ones ( Sarmiento-Gárces and Amat-García 2014 ). Distribution. This species is distributed in the Colombian Andes, in the three Andean cordilleras and is quite variable throughout its range, where the populations of the Eastern Cordillera are those that best match the description made by Felsche (1901) (see in Sarmiento- Gárces and Amat-García 2014). The species is found in Colombia between 500–2200 m a.s.l., consistent with the records of the species in the study area between 700– 1200 m a.s.l. This abundant species was found to prefer forested habitats (330 individuals) and was found also in wooded grasslands (two records).