<document id="0734C0D95D46883BC4730D23744C19DF" ID-DOI="10.5281/zenodo.3678492" ID-GBIF-Dataset="8e18452b-1d9d-462e-bac8-8c86eeb4ddc3" ID-ISSN="1942-1354" ID-Zenodo-Dep="3678492" ID-ZooBank="930EAB11-37FA-41B0-980A-1A4736527842" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582295086673" checkinUser="felipe" docAuthor="Hielkema, Auke J. &amp; Hielkema, Meindert A." docDate="2019" docId="03D7FF1CFFD7FFF1FF48FBB734DCFE0E" docLanguage="en" docName="InsectaMundi.0732.1-306.pdf.imf" docOrigin="Insecta Mundi 732 (732)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ataenius complicatus Harold 1869" docType="treatment" docVersion="12" lastPageNumber="27" masterDocId="FFEE8764FFCBFFEDFF88FB6E317CFFF0" masterDocTitle="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas" masterLastPageNumber="306" masterPageNumber="1" pageNumber="27" updateTime="1698721431862" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="EE386671A68CD3291E4E13F5A364F0CB">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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§ 
<taxonomicName id="4C7E3589FFD7FFF1FF5DFBB733CCFF01" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[213,688,217,241]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B90A9218FFD7FFF1FF5DFBB73080FF01" bold="true" box="[213,508,217,241]" italics="true" pageId="28" pageNumber="27">Ataenius complicatus</emphasis>
<bibRefCitation id="EFEF33FBFFD7FFF1FD8BFBB733CCFF01" author="Harold, E." box="[515,688,217,241]" journalOrPublisher="Coleopterologische Hefte" pageId="28" pageNumber="27" pagination="95 - 104" part="5" refId="ref206935" refString="Harold, E. von. 1869 a. Diagnosen neuer Coprophagen. Coleopterologische Hefte 5: 95 - 104." title="Diagnosen neuer Coprophagen" type="journal article" year="1869">Harold, 1869</bibRefCitation>
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<emphasis id="B90A9218FFD7FFF1FF60FA6E30A8FEE5" box="[232,468,256,277]" italics="true" pageId="28" pageNumber="27">Ataenius complicatus</emphasis>
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(
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- 
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)
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Distribution. 
<collectingCountry id="F3690E9AFFD7FFF1FEF2FA4830B2FECE" box="[378,462,294,318]" name="Brazil" pageId="28" pageNumber="27">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFFD7FFF1FDACFA483275FECE" author="Harold, E." box="[548,777,294,318]" journalOrPublisher="Coleopterologische Hefte" pageId="28" pageNumber="27" pagination="95 - 104" part="5" refId="ref206935" refString="Harold, E. von. 1869 a. Diagnosen neuer Coprophagen. Coleopterologische Hefte 5: 95 - 104." title="Diagnosen neuer Coprophagen" type="journal article" year="1869">Harold 1869a: 102</bibRefCitation>
; A. 
<bibRefCitation id="EFEF33FBFFD7FFF1FCBFFA483561FECE" author="Schmidt" box="[823,1053,294,318]" firstAuthor="Schmidt" pageId="28" pageNumber="27" pagination="1 - 111" refId="ref223873" refString="Schmidt, A. 1910 b. Aphodiinae. p. 1 - 111. In: S. Schenkling. Coleopterorum Catalogus Pars 20. W. Junk; Berlin. 111 p." type="book chapter" year="1910" yearSuffix="b">Schmidt 1910b: 73</bibRefCitation>
, 
<bibRefCitation id="EFEF33FBFFD7FFF1FBA2FA4835CDFECE" author="Schmidt" box="[1066,1201,294,318]" firstAuthor="Schmidt" pageId="28" pageNumber="27" refId="ref224130" refString="Schmidt, A. 1922 b. Coleoptera Aphodiinae. In: Das Tierreich. 45. Lieferung. Walter de Gruyter &amp; Co.; Berlin and Leipzig. 614 p." type="book" year="1922" yearSuffix="b">1922b: 425</bibRefCitation>
; 
<bibRefCitation id="EFEF33FBFFD7FFF1FB36FA483193FEAE" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="28" pageNumber="27" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 214</bibRefCitation>
; 
<bibRefCitation id="EFEF33FBFFD7FFF1FF73FA28308DFEAE" author="Cartwright, O. L." box="[251,497,326,350]" journalOrPublisher="The Coleopterists Bulletin" pageId="28" pageNumber="27" pagination="41 - 43" part="27" refId="ref196061" refString="Cartwright, O. L. 1973. Additional lectotype designations in the Aphodiinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 27 (1): 41 - 43." title="Additional lectotype designations in the Aphodiinae (Coleoptera: Scarabaeidae)" type="journal article" year="1973">Cartwright 1973: 42</bibRefCitation>
(designation 
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<emphasis id="B90A9218FFD7FFF1FCCDFA283536FEAE" box="[837,1098,326,350]" italics="true" pageId="28" pageNumber="27">Ataenius complicatus</emphasis>
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); 
<bibRefCitation id="EFEF33FBFFD7FFF1FBE9FA28342BFEAE" author="Stebnicka" box="[1121,1367,326,350]" firstAuthor="Stebnicka" pageId="28" pageNumber="27" pagination="1 - 2" refId="ref226727" refString="Stebnicka, Z. T. 2006 a. The New World species of Ataenius Harold, 1867. VII. Revision of the A. complicatus - group (Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia 49 B (1 - 2): 89 - 114." type="journal article" year="2006" yearSuffix="a">Stebnicka 2006a: 94</bibRefCitation>
(NA), 2007b: 53. 
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<collectingCountry id="F3690E9AFFD7FFF1FEB5FA0830B5FE8E" box="[317,457,358,382]" name="Venezuela" pageId="28" pageNumber="27">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFFD7FFF1FE5CFA0833BFFE8E" author="Stebnicka" box="[468,707,358,382]" firstAuthor="Stebnicka" pageId="28" pageNumber="27" pagination="1 - 2" refId="ref226727" refString="Stebnicka, Z. T. 2006 a. The New World species of Ataenius Harold, 1867. VII. Revision of the A. complicatus - group (Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia 49 B (1 - 2): 89 - 114." type="journal article" year="2006" yearSuffix="a">Stebnicka 2006a: 94</bibRefCitation>
, 
<bibRefCitation id="EFEF33FBFFD7FFF1FD45FA08323DFE8E" author="Stebnicka" box="[717,833,358,382]" firstAuthor="Stebnicka" pageId="28" pageNumber="27" refId="ref226889" refString="Stebnicka, Z. T. 2007 b. The genus Ataenius Harold, 1867 (Coleoptera: Scarabaeidae) of New World. Iconography. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences; Krakow, Poland. 155 p." type="book" year="2007" yearSuffix="b">2007b: 53</bibRefCitation>
. 
<emphasis id="B90A9218FFD7FFF1FCC2FA0832E3FE8E" bold="true" box="[842,927,358,382]" pageId="28" pageNumber="27">Other:</emphasis>
A. 
<bibRefCitation id="EFEF33FBFFD7FFF1FC4BFA0835E1FE8E" author="Schmidt" box="[963,1181,358,382]" firstAuthor="Schmidt" pageId="28" pageNumber="27" pagination="1 - 111" refId="ref223873" refString="Schmidt, A. 1910 b. Aphodiinae. p. 1 - 111. In: S. Schenkling. Coleopterorum Catalogus Pars 20. W. Junk; Berlin. 111 p." type="book chapter" year="1910" yearSuffix="b">Schmidt 1910b: 73</bibRefCitation>
(Guatemala, Mexico), 1922b: 425 (Guatemala, Mexico); 
<bibRefCitation id="EFEF33FBFFD7FFF1FDEFFAE832FFFE6E" author="Blackwelder, R. E." box="[615,899,390,414]" journalOrPublisher="United States National Museum Bulletin" pageId="28" pageNumber="27" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 214</bibRefCitation>
(Guatemala, Mexico); 
<bibRefCitation id="EFEF33FBFFD7FFF1FB2AFAE834DCFE6E" author="Stebnicka" box="[1186,1440,390,414]" firstAuthor="Stebnicka" pageId="28" pageNumber="27" pagination="1 - 2" refId="ref226727" refString="Stebnicka, Z. T. 2006 a. The New World species of Ataenius Harold, 1867. VII. Revision of the A. complicatus - group (Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia 49 B (1 - 2): 89 - 114." type="journal article" year="2006" yearSuffix="a">Stebnicka 2006a: 94</bibRefCitation>
(Argentina, Belize, Bolivia, Colombia, Costa Rica, Curaçao, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru), 2007b: 53 (Argentina, Belize, Bolivia, Colombia, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Netherlands Antilles, Nicaragua, Panama, Paraguay, Peru).
</paragraph>
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