Exploring the basal topology of the heteronemertean tree of life: establishment of a new family, along with turbotaxonomy of Valenciniidae (Nemertea: Pilidiophora: Heteronemertea) Author Kajihara, Hiroshi Author Abukawa, Shushi Author Chernyshev, Alexei V. text Zoological Journal of the Linnean Society 2022 196 503 548 journal article 133817 10.1093/zoolinnean/zlac015 372f8265-c53c-43fa-9448-07d5a58d311f 0024-4082 7037958 78C56409-FCCF-4116-8D8C-FF66B247C56C GENUS BASEODISCUS DIESING, 1850 Polia Delle Chiaje, 1822 in Delle Chiaje (1822–29) (see Remarks below); not Polia Ochsenheimer, 1816: 73 (Lepidoptera) . Baseodiscus Diesing, 1850: 243 ; type species Polia delineata Delle Chiaje, 1822 , fixed by monotypy. Taeniosoma Stimpson, 1857: 162 ; type species not fixed (either Taeniosoma aequale Stimpson, 1857 or Taeniosoma septemlineatum Stimpson, 1857 ). Eupolia Hubrecht, 1887: 10–11 ; type species Polia delineata Delle Chiaje, 1822 , fixed by original designation. Balanocephalus Kennel, 1891: 282 ; type species Balanocephalus pellucidus Kennel, 1891 , fixed by monotypy. Nematodemus Graff, 1899: 513 ; type species Nematodemus lumbricoides Graff, 1899 , fixed by monotypy. Diagnosis: Given by Gibson (1994: 114) . An additional feature is the proboscis musculature consisting of inner circular, middle diagonal, and outer longitudinal layers ( Magarlamov & Chernyshev, 2011 ). Remarks: The publication year of the generic name Polia Delle Chiaje is 1822 rather than 1825 (e.g. Gibson, 1995: 475 ) or 1827 (e.g. Neave, 1940: 844 ) as previously regarded, in accordance with Article 12.2.7 of the Code ( ICZN, 1999 ); the type species has not been fixed. The name Polia Delle Chiaje was first used for the following six nominal species, originally appearing as captions for plates supposedly published in 1822, while the written descriptions for these species were supplied in Volume 2 (issued in 1825 or 1827) (see Remarks for Baseodiscus delineatus below) or Volume 3 (issued in 1828): Polia caerulescens Delle Chiaje, 1822 (pl. XLIII, fig. 9) (description in Volume 3 as ‘ P. cilestra ’: 172–173); Polia delineata Delle Chiaje, 1822 (pl. XXVIII, fig. 4) (description in Volume 2 as ‘ P. lineata ’ on p. 409 and as ‘ P. delineata ’ on pp. 427–428; the specific name also appears on p. 444 in the explanations for plates); Polia geniculata Delle Chiaje, 1822 (pl. XLIII, fig. 10; LXXVIII, figs 5, 6) (description in Volume 3: 177); Polia oculata Delle Chiaje, 1822 (LXXVIII, fig. 8) (description in Volume 3: 172, 177; the name also occurs on p. 181 in the explanations for plates); Polia punctata Delle Chiaje, 1822 (pl. LXVIII, figs 7, 11) (description in Volume 3, as ‘ P. punteggiata ’ on p. 172 and ‘ P. punctata ’ on p. 177; the name also occurs on p. 181 in the explanations for plates); and Polia siphunculus Delle Chiaje, 1822 (pl. XXVIII, figs 1–3) (description in Volume 2, as ‘ P. sipuncolo ’ on pp.406–408 and ‘ P. sipunculus ’ on p. 427). However, these complications do not overturn precedence of the senior homonym Polia Ochsenheimer, 1816 ( Lepidoptera : Noctulidae) over Polia Delle Chiaje, 1822 .