<document id="624D9A8E89655FAE44C3109C6CA9F3FB" ID-DOI="10.3897/zookeys.1230.136906" ID-ZooBank="0FBF41B0-1D35-4A31-9653-B1F475FBDF80" ID-publisher-id="136906" URI-arpha="3B6146E9-7C33-583C-A633-97ED3C18A10A" URI-zoobank="http://zoobank.org/0FBF41B0-1D35-4A31-9653-B1F475FBDF80" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1741198404937" checkinUser="pensoft" docAuthor="Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J. & Liew, Thor-Seng" docDate="2025" docId="58EA9368F87952DA8C2C4FEDA56DD2BC" docLanguage="en" docName="ZooKeys 1230: 99-154" docOrigin="ZooKeys 1230" docSource="https://zookeys.pensoft.net/article/136906/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Microcystina undetermined-5" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="154" masterDocId="3B6146E97C33583CA63397ED3C18A10A" masterDocTitle="From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia" masterLastPageNumber="154" masterPageNumber="99" pageNumber="99" updateTime="1741227862456" updateUser="ExternalLinkService">
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<mods:title id="2970B63E0481B364E1C13A3C86480745">From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia</mods:title>
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<mods:namePart id="0A14A6D16836EDFCA5A0B89CD78A36D9">Goh, Soo-Mun</mods:namePart>
<mods:affiliation id="33B636D71B36EF95D41695FD5FF03EEF">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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<mods:namePart id="F20A96B33F2206F395F22214BA484BF2">Jasrul, Dulipat</mods:namePart>
<mods:affiliation id="73EDF9356F75BD7D56202ADAC52E29EF">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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<mods:namePart id="F390021DE5C058FF0CCA9A215545905E">Lee, Mei-Yi</mods:namePart>
<mods:nameIdentifier id="20DD8461128AE420D6FC4F259F026E2F" type="ORCID">https://orcid.org/0009-0005-8200-3508</mods:nameIdentifier>
<mods:affiliation id="35986359FF14FC50E0BCDCAAB28EC77F">School of Biological Science, Universiti Sains Malaysia, Penang, Malaysia</mods:affiliation>
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<mods:namePart id="BF882A573CE7A60714B3B84AB67419F6">Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation id="45C34BFF9EE94C0027B5D7A57590EF9A">jk. artandscience, Lauwerbes 8, 2318 AT, Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="68A7A728D96FAA691254E55C1F5760E2">Liew, Thor-Seng</mods:namePart>
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<mods:affiliation id="894F9BE5CEA722C20099FD23976D6C0E">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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<treatment id="58EA9368F87952DA8C2C4FEDA56DD2BC" ID-DOI="http://doi.org/10.5281/zenodo.14977472" ID-Zenodo-Dep="14977472" ID-arpha="58EA9368-F879-52DA-8C2C-4FEDA56DD2BC" LSID="urn:lsid:plazi:treatment:58EA9368F87952DA8C2C4FEDA56DD2BC" httpUri="http://treatment.plazi.org/id/58EA9368F87952DA8C2C4FEDA56DD2BC" scope_class="Gastropoda" scope_family="Ariophantidae" scope_phylum="Mollusca" scope_subClass="Heterobranchia" scope_superFamily="Helicarionoidea">
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<paragraph id="15360881B74F182C7EC9C51A9489E1EF">
<taxonomicName id="BD234F298EE12D8EEC044EBC7D0C7404" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="undetermined-5">
<emphasis id="55679AB668BDB2D1B1963C89F0E34C0E" italics="true">Microcystina</emphasis>
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sp. 5
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<paragraph id="5F6D7E29DBB6CA011D427209E7BC90CB">
<figureCitation id="C2250E59CBE56FC02C03EF2140E11B8D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Microcystina Penang sp. 2 BOR / MOL 15458 B Microcystina Penang sp. 4 BOR / MOL 15755 C Microcystina Penang sp. 5 BOR / MOL 15762 D Microcystina Penang sp. 6 BOR / MOL 15756. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1230.136906.figure9" httpUri="https://binary.pensoft.net/fig/1270444">Fig. 9 C</figureCitation>
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<paragraph id="6A4264468A2C95F4422EC40A3A512B41">
<heading id="8F86CD69FB967E8F41F875580657F10B" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="745B5B286380FD3AAABA07E2E7B781D1">
<materialsCitation id="5B69E2B9FDD18B58E5BFB2D231662D0E" collectingDate="2022-10-25" collectionCode="T" collectorName="T. S. Liew & J. Dulipat" country="Malaysia" county="Penang Hill" elevation="290" latitude="5.3977" location="Forest near Dam, Plot. 26" longLatPrecision="7" longitude="100.2612" municipality="Air Itam" specimenCode="BOR/MOL 15464" specimenCount="1" stateProvince="Pulau Pinang">
<collectingCountry id="4871E1D308CF61E20EA6E9B11B250055" name="Malaysia">Malaysia</collectingCountry>
•
<collectingCounty id="90937F3A9E3AB19DEE78E392DBD79B9F">Penang Hill</collectingCounty>
,
<collectingMunicipality id="FAD73EE7664E2DEA6512AFE76BADCFB7">Air Itam</collectingMunicipality>
,
<location id="52C4588B148B04FD088C4D62F4DE6C1D" LSID="urn:lsid:plazi:treatment:58EA9368F87952DA8C2C4FEDA56DD2BC:52C4588B148B04FD088C4D62F4DE6C1D" country="Malaysia" county="Penang Hill" latitude="5.3977" longLatPrecision="7" longitude="100.2612" municipality="Air Itam" name="Forest near Dam, Plot. 26" stateProvince="Pulau Pinang">Forest near Dam, Plot. 26</location>
;
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<geoCoordinate id="76A1F07DC7EA28F8E4D0BE6C39AB3E0D" degrees="5.3977" direction="north" orientation="latitude" precision="5" value="5.3977">5.3977 ° N</geoCoordinate>
,
<geoCoordinate id="C827EDBDD5BB2A370565CCCC76DE2E4A" degrees="100.2612" direction="east" orientation="longitude" precision="5" value="100.2612">100.2612 ° E</geoCoordinate>
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;
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<quantity id="A95D92853E1C6FE08A4256DD4BBE5DD1" metricMagnitude="2" metricUnit="m" metricValue="2.9" unit="m" value="290.0">290 m</quantity>
a. s. l.
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;
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<collectingDate id="2FB7F78E555C848A285186D7399C8F2A" value="2022-10-25">25 Oct. 2022</collectingDate>
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;
<collectorName id="C6F274F245A3D52681A3F2994446F96C">T. S. Liew</collectorName>
,
<collectorName id="56FB45366C5755E4ABFBB0E7ECC2C6A9">J. Dulipat</collectorName>
leg.;
<specimenCode id="FF101EC9C10723136FFA6DFD17463712">BOR/MOL 15464</specimenCode>
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•
<materialsCitation id="5F35C8B523E4BF352FE3255378EDE183" collectingDate="2023-02-04" collectionCode="T" collectorName="T. S. Liew & A. Yusni" country="Malaysia" county="Penang Hill" elevation="610" latitude="5.421" location="By Path H, Plot. 33" longLatPrecision="7" longitude="100.2701" specimenCode="BOR/MOL 15762" specimenCount="1" stateProvince="Pulau Pinang">
<collectingCounty id="6DABA0D2EC9BA4B2861F638854063601">Penang Hill</collectingCounty>
,
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;
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<geoCoordinate id="69BB5B8397F780F4F169367595B77A62" degrees="5.4210" direction="north" orientation="latitude" precision="5" value="5.421">5.4210 ° N</geoCoordinate>
,
<geoCoordinate id="ABD7A30A88691CBBF1D26CD4B4B3C851" degrees="100.2701" direction="east" orientation="longitude" precision="5" value="100.2701">100.2701 ° E</geoCoordinate>
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;
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<quantity id="5E748948E1DD9459DFAE0E14F5DB5D58" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">610 m</quantity>
a. s. l.
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;
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<collectingDate id="C4C2EB3F2DE4DC2BE40A600E22F0AC1C" value="2023-02-04">4 Feb. 2023</collectingDate>
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;
<collectorName id="B04CAF01B06FB957FBE59B8C7CDE31EB">T. S. Liew</collectorName>
,
<collectorName id="1E531A365B96D87B9E9D3B80111BA681">A. Yusni</collectorName>
leg.;
<specimenCode id="D7222AE2EEB2CCC450A95FA87FEBE092">BOR/MOL 15762</specimenCode>
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.
</paragraph>
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<subSubSection id="SECID0EOZDI" type="remarks">
<paragraph id="6E7A79CDA81AC47BD08CAE42DF2D2100">
<heading id="E057468097AFB65858B45F1BB204B2A9" reason="title">Remarks.</heading>
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<paragraph id="7F65578C64F0E073237F9A203915C243">
This is a new record for
<collectingCounty id="A8BB7A8B30C38AAD63D5C6648E39C06E">
<collectingRegion id="10B576E88C74B2ABD4538A8B4FD98B02" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
Hill
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. The shell spire is hardly raised, with a height less than half of the last whorl height. On the upper shell, there are densely spirally arranged pits on the apical and subsequent whorls. This species is unique among the
<taxonomicName id="9F965BB4871907E41A665F8BEEDDB733">
<emphasis id="04302F26500BD26885DE5D7F30BD5A87" italics="true">Microcystina</emphasis>
species
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from
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<collectingRegion id="3C9F2F7E6192B14CD5F66C2CFBC648F5" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
Hill
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in having densely spirally arranged pits on the whorls near the periphery. Below the shell, there are densely spirally arranged pits that continue from the upper shell and fade away towards the umbilicus.
</paragraph>
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</treatment>
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