Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae) Author Selnekovič, Dávid D930D757-D929-4248-A29C-A857E0505345 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. david.selnekovic@uniba.sk Author Jäch, Manfred A. D749707A-8823-4110-8D51-BF5AF4E2820F Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria. manfred.jaech@nhm-wien.ac.at Author Kodada, Ján 6E88BFBB-8769-44EC-8285-29E357CEC064 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. jan.kodada@uniba.sk text European Journal of Taxonomy 2024 2024-09-11 957 1 229 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247 journal article 10.5852/ejt.2024.957.2651 2118-9773 13773385 9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3 Elmomorphus punctulatus sp. nov. urn:lsid:zoobank.org:act: D8306DD8-ED3F-47D2-82BD-9B9FB26A9D76 Figs 58–59 , 60B , 112C Differential diagnosis Elmomorphus punctulatus sp. nov. ( Fig. 58 ) belongs to a group of species characterised by having the dorsal plastron distributed on the anterior portions of the head and the posterolateral portions of the elytra. Elytral punctures are scattered over the entire surface, and striae are absent. Within this group, E. nepalensis and E. punctulatus may be recognised by the absence of long and erect setae or setal clusters on the labrum, prosternal process, or metaventrite of males. The median part of the metaventrite is weakly convex in males, whereas in other species it is depressed. The tibiae are simple in both sexes, not expanded, and without teeth. The species is further characterised by the strongly convex body ( Fig. 58 ), distinctly rounded lateral sides of the pronotum, the shape of the aedeagus ( Fig. 59 ), and the presence of large separate spines on the bursa copulatrix ( Fig. 60B ). Differences between E. punctulatus and E. nepalensis are listed under the latter species. Etymology The epithet is a Latin adjective in the nominative singular meaning ‘punctulate’, referring to the relatively dense punctation on pronotum and elytra. Type material Holotype CHINA – Yunnan Province • ♂ ; “ CHINA : Yünnan , Xishuangbanna ca. 6km NW Mengla 8.11.1999 , ca. 700 m leg. Jäch , et al. ( CWBS 365 )”; IAECAS . Paratypes CHINA – Yunnan Province • 1 ♂ , 1 ♀ ; “ CHINA : Yünnan , Xishuangbanna ca. 6 km NW Mengla 9.11.1999 , ca. 700 m leg. Jäch , et al. ( CWBS 367 )”; CKB , NMW • 1 ♂ , 1 ♀ ; “ CHINA : Yünnan , Xishuangbanna ca. 20km W Jinghong 11.11.1999 , ca. 1000 m leg. Jäch , et al. (374)”; NMW . Type locality China , Yunnan Province , Xishuangbanna Dai Autonomous Prefecture, Mengla County, ca 6 km northwest of Mengla, ca 700 m a.s.l.; small stream, ca 0.5 m wide, flowing through primary forest ( CWBS 365; Jäch & Ji 2003 ). Description Measurements (mm): TL: ♂♂ 2.55–2.68 (n =3), ♀♀ 2.65–2.95 (n= 2); PL: ♂♂ 0.63–0.73 (n=3), ♀♀ 0.70–0.74 (n =2); PW: ♂♂ 1.20–1.24 (n= 3), ♀♀ 1.32–1.35 (n =2); EL: ♂♂ 1.92–1.95 (n=3), ♀♀ 1.95–2.21 (n=2); EW: ♂♂ 1.35–1.43 (n= 3), ♀♀ 1.46–1.53 (n=2); PhL: 0.51–0.52 (n =2); PrL: 0.34– 0.35 (n =2). Fig. 58. Elmomorphus punctulatus sp. nov. , paratype, male from China, locality CWBS 367 (CKB), TL: 2.63 mm. Fig. 59. Elmomorphus punctulatus sp. nov. , holotype (IAECAS), aedeagus. A . Lateral aspect. B . Ventral aspect. Scale bar: 0.1 mm. Body ( Fig. 58 ) oval, strongly convex dorsally. Integument brown to black; trochanters, tarsi, antennae, and mouthparts paler. Pubescence consisting of very short yellowish setae. Plastron present on anterior portions of head, apicolateral portions of elytra, and on ventral surface except for prosternal process, median part of metaventrite, and middle of ventrites 1–2. Head with round setiferous punctures slightly smaller than an eye facet, separated by 0.5–1.0 × puncture diameter; surface between punctures smooth; plastron on frontoclypeus and anterolateral portions of vertex. Labrum microreticulate, with small setiferous punctures; length of setae equal in both sexes. Anterior margin of clypeus straight. Antennae 9-segmented. Eyes oval, interfacetal setae short, ID: ♂♂ 0.46–0.49 mm (n =3), ♀♀ 0.48–0.52 mm (n=2), APD/ID: ♂♂ 1.49–1.58 (n=3), ♀♀ 1.56 (n =2). Pronotum wider than long, widest at base, strongly convex, PW/PL: ♂♂ 1.69–1.90 (n =3), ♀♀ 1.82–1.88 (n=2); surface smooth, with round setiferous punctures; plastron absent; rim of anterior margin as wide as three eye facets, interrupted in middle; lateral pronotal sides convergent, distinctly rounded in anterior half. Prosternal process with lateral edges divergent, straight, posterior edge broadly V-shaped; lateral portions raised, long setae absent in both sexes; median keel slightly arcuate. Scutellum approximately as long as wide, smooth, with round punctures. Metaventrite slightly convex in median part; groups of long setae missing in both sexes. Elytra oval, widest around middle; EL/EW: ♂♂ 1.36–1.42 (n =3), ♀♀ 1.33–1.45 (n=2); surface with fine microreticulation and dense setiferous punctures; plastron covering apicolateral portions. Tibiae slightly curved, not expanded, teeth absent; PrTL/PL: ♂♂ 0.83–0.96 (n=3), ♀♀ 0.87–0.91 (n= 2). Terminal protarsomere as long as three preceding tarsomeres combined. Fig. 60. Bursa copulatrix and vagina. A . Elmomorphus nepalensis Satô, 1981 , female from Nepal (NMW). B . E. punctulatus sp. nov. , paratype from China, locality CWBS 367 (NMW). Scale bar: 0.1 mm. Ventrites covered with plastron, except in middle of ventrites 1–2. Ventrite 5 with very short, rather indistinct longitudinal keel at apex; in females with small triangular excision. Aedeagus ( Fig. 59 ): phallobase rather long and slender, slightly expanded proximally, PhL/PrL: 1.47–1.48 (n= 2), basal opening rather large; parameres regularly narrowed to apices, slightly curved ventrad, apices narrowly rounded (lateral aspect); penis narrowly rounded apically. Ovipositor: valvifers long; coxites asymmetrical, right one ca 1.30 × as long as left one. Bursa copulatrix with moderately large, separated spines in dorsal and lateral portions ( Fig. 60B ); spermatheca tubular. Secondary sexual dimorphism Ventrite 5 in females with small triangular apical excision. Distribution China ( Yunnan ) ( Fig. 112C ).