Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae) Author Selnekovič, Dávid D930D757-D929-4248-A29C-A857E0505345 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. david.selnekovic@uniba.sk Author Jäch, Manfred A. D749707A-8823-4110-8D51-BF5AF4E2820F Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria. manfred.jaech@nhm-wien.ac.at Author Kodada, Ján 6E88BFBB-8769-44EC-8285-29E357CEC064 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. jan.kodada@uniba.sk text European Journal of Taxonomy 2024 2024-09-11 957 1 229 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247 journal article 10.5852/ejt.2024.957.2651 2118-9773 13773385 9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3 Elmomorphus paramontanus Kodada, Selnekovič & Jäch sp. nov. urn:lsid:zoobank.org:act: BCB85417-4F7A-4DA9-A83A-9C0639A59A91 Figs 1B–C, E–G , 2F–J , 3E , 4A, F, I , 40–41 , 107C–D , 111A Differential diagnosis Elmomorphus paramontanus sp. nov. ( Fig. 40 ) belongs to a group of species characterised by having a plastron on the dorsal cranial surface (except for a triangular area on the vertex), on the anterior angles of the pronotum, and the lateral sides of the elytra. Elytral punctures scattered over the entire surface, striae absent. Within this group, E. corpulentus sp. nov. , E. ellipticus sp. nov. , E. montanus , E. paramontanus , and E. prosternalis may be recognised by the pronotal plastron being confined to the anterior angles. The differences between E. paramontanus , E. corpulentus , and E. ellipticus are listed under the latter two species. Elmomorphus montanus differs from E. paramontanus in the parameres being shorter in relation to the phallobase, PhL/PrL: 2.30 in E. montanus versus PhL/PrL: 1.37–1.81 (1.57 ± 0.11, n=25) in E. paramontanus ( Fig. 41 ). Fig. 40. Elmomorphus paramontanus Kodada, Selnekovič & Jäch sp. nov. , paratype, male from type locality (NMW), TL: 3.10 mm. Fig. 41. Elmomorphus paramontanus , Kodada, Selnekovič & Jäch sp. nov. , paratype from Thailand (CKB), aedeagus. A . Lateral aspect. B . Ventral aspect, detail of parameres and penis. C . Ventral aspect. Scale bars: 0.1 mm. Etymology The epithet is a Latin adjective in the nominative singular referring to the overall similarity with E. montanus . Type material Holotype THAILAND – Mae Hong Son Province • ♂ ; “Thai 1-8.V.1992 Ban-si Lang Mae Hong Son J.Horák leg.”; NMW . Paratypes CHINA – Hong Kong • 2 ex. ; “ HONGKONG University Campus at light , 1996 leg. G. de Rougemont ”; NMW . – Yunnan Province • 2 ♂♂ ; “CHINA:Yünnan, Xishuangbanna ca. 15 km W Menglun , 5.11.1999 , ca. 700 m leg. Jäch , et al. (CWBS 355)”; NMW • 3 ex. ; “ CHINA : Yünnan , Xishuangbanna ca 10km NW Menglun 7.11.1999 , ca. 700 m leg. Jäch , et al. (CWBS 359)”; NMW • 12 ♂♂ , 14 ♀♀ , 44 ex. ; “ CHINA : Yünnan , Xishuangbanna , ca. 50 km SSE Menglun , Mengyuan , 6.11.1999 , ca. 700 m , leg. Jäch et al. (CWBS 361)”; NMW • 4 ♀♀ , 6 ex. ; “ CHINA : Yünnan , Xishuangbanna ca. 6km NW Mengla 8.11.1999 , ca. 700 m , leg. Jäch et al. (CWBS 364)”; NMW • 1 ex. ; “ CHINA : Yünnan , Xishuangbanna ca. 50km NW Mengla 9.11.1999 , ca. 800 m leg. Wang & Wei (CWBS 370)”; NMW • 7 ex. ; “ CHINA : Yünnan , Xishuangbanna ca. 7km NE Jinghong 12.11.1999 , ca. 600 m leg. Jäch , et al. (CWBS 377)”; NMW • 12 ex. ; “ CHINA : S-YUNNAN ( Xishuangbanna ) 23 km NW Jinghong Na Ban Village (NNNR [ Nabanhe National Nature Reserve ]) | N22°10.04 E100°39.52 680m 05.VI.2008 LF [ light trap ] leg. A. Weigel station”; CWW • 7 ex. ; same data as for preceding, but labels written in different style; CWW • 1 ex. ; “ CHINA : S-YUNNAN ( Xishuangbanna ), 20km NW Jinghong , Man Dan (NNNR [ Nabanhe National Nature Reserve ]), N 22°07″80′ , E 100°40″05′ [minutes and seconds interchanged], 720m , 23.V. 2008 , leg. A.Weigel , LF [ light trap ]”; CWW . – Guangdong Province • 1 ex. ; “ CHINA : Guangdong Prov., 35 km NE Huaiji 24°00′25″N 112°26′09″E 3.11.2001 , ca. 130 m Jäch & Komarek (CWBS 463)”; NMW • 1 ex. ; “ CHINA : Guangdong Prov . 40 km ENE Zengcheng 23°17′12″N 114°03′42″E 11.11.2001 , ca. 140 m Komarek & Wang (CWBS 492)”; NMW . LAOS – Houa Phan Province • 1 ex. ; “LAOS-NE, Houa Phan prov., 20°13′09-19″N 103°59′54″- 104°00′03″E , 1480-1510m , PHOU PANE Mt. , 22.iv.-14.v.2008 , Vít Kubáň leg.”; NMW . – Luang Namtha Province • 4 ex. ; “N-LAOS: Prov. Lg. Nam Tha ca 20km NE Muang Sing 10.6.1996 , 700m leg. Schillhammer (20)”; NMW • 58 ex. ; “N-LAOS: Prov. Lg. Nam Tha ca. 5 km Muang Sing 650 m Huay Giulom riv., 10.6.1996 leg. Schillhammer (21)”; NMW • 4 ex. ; “N-Laos: Prov. Lg. Nam Tha , 5-10 km SW Muang Sing , 11.6.1996 , 600m , leg. Schillhammer (23)”; NMW • 2 ex. ; “N-LAOS: Prov. Lg. Nam Tha ca. 20km SE Muang Sing 12./ 13.6.1996 , 950m leg. Schillhammer (25)”; NMW • 40 ex. ; “N-Laos, Prov. Lg. Nam Tha , ca. 30 km NW Lg. Nam Tha , 16./ 18.6.1996 , 800 m , leg. Schillhammer (28, 30)”; NMW • 20 ex. ; “ LAOS north, 24.–30.V 1997 20 km NW Louang Namtha N 21°09.2 , E 101°18.7 , alt. 900± 100 m E. Jendek & O. Šauša leg.”; CKB . – Luang Prabang Province • 3 ex. ; “ IX. 1992 , 250 m , 10 km N Luang Prabang Mekong-Ufer [banks], Licht [at light], leg. Insomsay Somsy ”; NMW . – Vientiane Province • 13 ex. ; “ Ban Pahom 24km N of Vang Vieng Vientiane Prov., LAOS 9. V. 2002 H. Yoshitomi leg.”; NMW . MALAYSIA – Perak State • 5 ex. ; “MALAYSIA 3.II.1992 PERAK: ca. 10 km NÖ [NE] Gerik leg. Jäch (24)”; NMW • 2 ex. ; “ MALAYSIA 4.II.1992 PERAK : Lawin, S Gerik leg. Jäch (25)”; NMW • 1 ex. ; “ MALAYSIA 4.II.1992 PERAK : Ö [E] Padang Gerus leg. Jäch (26)”; NMW . MYANMAR • 3 ex. ; “MYANMAR N ( Burma ) 21 km E Putao , H- 550m Nan Sa Bon vill 01.- 05.05.1998 leg. S. Murzin & V. Siniaev ”; NMW • 21 ex. ; “ MYANMAR : Kachin State 3.8km SW Putao , 30.5./ 1.6.1999 Ma Kyaw Wa riv., ca. 480m 27°18.90′N 97°22.93′E leg. Schillhammer & Schuh (59)”; NMW • 5 ex. ; “ MYANMAR : Kachin State 12km S Putao , 31.5.1999 W Mularshidi vill., 500 - 550m 27°15.13′N 97°24.95′E leg. Schillhammer & Schuh (60)”; NMW • 2 ex. ; “ MYANMAR : Kachin State ca. 12km S Putao , 2.6.1999 W Mularshidi vill., 50 - 550m 27°14.98′N 97°24.40′E leg. Schillhammer & Schuh (61)”; NMW . THAILAND – Mae Hong Son Province • 5 ♂♂ , 9 ♀♀ , 10 ex. ; same collection data as for holotype; CKB , NMW • 17 ex. ; “NW THAI 23.-31.V. MAE HONG SON 1200m Ban Si Lang 1991 J. Horák leg.”; CKB • 12 ex. ; “ THAILAND : 13.11.1995 3 km SE Mae Hong Son leg. Zettel (14b)”; NMW • 6 ♂♂ , 9 ♀♀ , 11 ex. ; “NW-Thai 23.-31. V. 1991 , Ban Si Lang , Mae Hong Son 1200m , J. Horák leg.”; CKB , NMW • 1 ♂ , 2 ♀♀ , 3 ex. ; “NW Thailand , 19.19N , 97.59E Mae Hong Son , 1991 Ban Si Lang , 1200m 23.-31.5., L.Dembický leg.”; NMW • 6 ex. ; “NW THAI. 23.-31.V. MAE HONG SON 1200m BAN SI LANG, 1991 ǀ COLL. BOUKAL”; CBB • 3 ♂♂ , 2 ♀♀ ; “NW-Thailand, 1.-7. V. 1992 , Mae Hong Son , Ban Si Lang , 1000 m , S. Bílý leg.”; NMW • 18 ex. ; “N-THAILAND, V.1992 Mae Hong Son Huai Sua Tao leg. Dembicky”; NMW • 1 ex. ; “NW THAILAND Mae Hong Son Suan Pu, 5.5.1992 Pacholatko & Dembicky ”; NMW • 3 ex. ; “ THAILAND : 11.11.1995 17km N Mae Hong Son Mok Cham Pae leg. Zettel (12a)”; NMW . – Chiang Mai Province • 24 ex. ; “ THAILAND 1989 Chiang Mai Malicky 10.-17.IV. ”; NMW • 2 ♂♂ , 1 ♀ ; “NW Thailand CHON [CHOM] THONG 24-27.IV.1991 J. Horak lgt.”; NMW • 1 ex. ; “NW THAI 24.-27.IV. CHON THONG 1991 J. Horák leg.”; CKB • 1 ♀ ; “NW-Thailand, 1991, Chon [ Chom ] Thong , 24. - 27.4., 18°26′N 98°41′E , L. Dembický leg.”; NMW • 1 ex. ; “N-THAILAND: Ch. Mai Chom Thong 24.- 26.4.1991 leg. Pacholatko”; NMW • 2 ♂♂ , 3 ♀♀ ; “N THAI, 15.-21.1. CHANG-MAI [ Chiang Mai ] env. 1992 Meo-Sa waterfalls M. Holecová leg.”; CKB • 28 ex. ; “ Thailand : Chiang Mai Suanrim, 25.III.1994 , Nam Mae Si ,WDS-A-1046 William D. Shepard , leg. | Collection W.D. Shepard ”; CSS • 2 ♀♀ ; “ Thailand 91 “Thanon Thong Chai”, D. Král & V. Kubáň , Thai , 10. - 16. V. 1991 , Chiang Dao 600 m , 19°24′N 98°55′E , Vít Kubáň leg.”; NMW • 5 ♂♂ , 8 ♀♀ ; “ Thailand 91 “Thanon Thong Chai”, D. Král & V. Kubáň , Thai, 26. - 28. V. 1991 , Palong 750 m , 19°55′N 99°06′E , Vít Kubáň leg.”; NMW • 2 ♂♂ , 1 ex. ; “ Thailand 91 “Thanon Thong Chai”, D. Král & V. Kubáň , Thai , 9. - 14. V. 1991 , Chiang Dao 350 m , 19°22′N 98°57′E , Vít Kubáň leg.”; NMW • 1 ♂ ; “ THAILAND 31.12. Huay Mae Sai Wf [waterfall] leg. Madl 1987”; NMW • 4 ♂♂ , 3 ♀♀ , 44 ex. ; “ North Thai , Maeo Khun Klang, 1350 m , Doi Inthanon, 16. X. 1983 , M Sakai”; NMW • 3 ♂♂ , 21 ex. ; “ Thailand , Maeo Khun Klang, 1300 m , Doi Inthanon, 17. X. 1983 , M. Tomokuni , at light”; NMW • 10 ex. ; “N-THAILAND: Chiang Mai 12 km W Chom Thong Mae Klang riv. 27.12. 1998 , 550 m , leg. P. Mazzoldi (7)”; NMW • 2 ex. ; “ Thailand : Doi Inthanon Mae Klang , 11.4.1996 , 18°32′N 98°36′E , 540 m , WRC 17, leg. Malicky ”; NMW • 4 ex. ; “NW THAIL. 15.-23.5. Chiang Mai Doi Inthanon 1989 leg. Malicky ”; NMW • 4 ♂♂ , 14 ex. ; “ Fang Thailand Nov. , 22. 1968, M. Sato leg.”; NMW • 24 ex. ; “ THAILAND 1989 Chiang Mai Malicky 10.-17.IV. ”; NMW • 1 ex. ; “NW-Thail.: Chiang Mai , 38°57′E 18°49′N , Zoo, 24.- 31.10.1988 Malicky & Chantaramon[g]kol LF [ light trap ]”; NMW • 18 ex. ; “N-THAILAND, 10.- 17.4. 18°49′N 98°57′E Chiang Mai , Zoo (Licht) [light] leg. Chantaramongkol & Malicky 1989”; NMW • 1 ex. ; “ THAILAND Phuphing Palace, 1400m , Doi Suthep-Pui, C. Chiang Mai , 8 Sep 1987 T. Yamasaki ”; NMW • 86 ex. ; “N-THAILAND NW Chiang Mai Pai City, 29.4.1993 Pacholatko & Dembicky ”; NMW • 1 ex. ; “N-THAILAND, Chiang M. Soppong - Pai 1.- 8.5.1993 , 1800m Pacholatko & Dembicky ”; NMW • 6 ex. ; “ THAILAND : 30./31.11. Chiang Mai Prov. 1995 W Mae Rim, Mae Sa NP leg. Zettel (2)”; NMW • 45 ex. ; “ THAILAND : 8.11.1995 Chiang Mai Prov. Chiang Dao , 500m leg. Zettel (10)”; NMW • 4 ex. ; “N-THAILAND Ban Sanpakia, 19°19′N 96°50′E 1000 m a.s.l. , 3.V.1996 , V. Kubáň leg.”; CKB . – Kanchanaburi Province • 3 ex. ; “ THAILAND : 5.4.1989 , Tham Than Lod NP [= Chaloem Rattanakosin National Park ], 500m 99°20′E 14°46′N , Malicky & Wanleelag”; NMW . – Nan Province • 1 ex. ; “N-THAILAND: Nan, 2.1.1999 Doi Phu Ka NP, 1400 m 1 km after Park HQ leg. P. Mazzoldi (24)”; NMW . – Nakhon Si Thammarat Province • 1 ex. ; “P. THAILAND Khao Luang N.P.-H.Q. 450m Nop Pi Tam , C. Nakhon Si Thammarat, 6 Aug 1987 , T. Yamasaki ”; NMW . – Phetchabun Province • 1 ex. ; “N-THAILAND, Nan Lom Sak - Dan Sai 17.- 19.6.1993 Pacholatko & Dembicky ”; NMW . – Phrae Province • 2 ex. ; “ THAILAND : 17./ 18.11.1995 Phrae Prov.Huai Kaet 50km NE Phrae leg. Zettel (17b)”; NMW . – Songkhla Province • 11 ex. ; “S-Thailand 600m ob [above] Ton Nga Chang WF [waterfall] 6°58′N , 100°12′E lg. Malicky, 4.-5.5.[19]93”; NMW . – Tak Province • 7 ♂♂ , 8 ♀♀ , 7 ex. ; “ Thailand 91 “Thanon Thong Chai”, D. Král & V. Kubáň , Thai, 26. IV. - 6. V. 1991 , Umphang 550 m , 16°04′N 98°53′E , Vít Kubáň leg.”; CKB , NMW . – Yala Province • 121 ♂♂ , 38 ♀♀ , 169 ex. ; “S-THAILAND 25.3.-22.4. BETONG YALA distr. 1993 leg. J.Horák & J.Strnad ”; CKB , NMW . VIETNAM – Bac Kan Province • 2 ex. ; “N-VIETNAM Bac Kan Prov., Ba Bè NP, (entry), 16.- 20.V.2014 22°25′07″, E 105°38′09″E 180-220m , leg. A. Skale ”; NMW . – Cao Bang Province • 1 ex. ; “N-VIETNAM Cao Bang Prov., vic. Tinh Tuc , Son Dong , Nui Pia Oac Nature Reserve , 9.-15.V.2014 22°37′55″N 105°52′98″E 860-1300m leg. A. Skale ”; NMW • 2 ex. ; “N- VIETNAM , Cao Bang Pr. vic. Tinh Tuc , Son Dong Nui Pia Oac Nature Res. 22°37′5″N , 105°53′20″ 11.V.2014 , 1000m leg. A. Weigel , by light”; CWW . – Lam Dong Province • 1 ♂ , 1 ♀ ; “S-Vietnam: 17.-21.4., 12 km N Dalat , 1995, Lang Bian | 12°03′N 108°27′E , 1580-1750 m , Pacholatko & Dembicky”; NMW • 56 ex. ; “S-VIETNAM: 22.- 24.4. 15km SW Bao Loc 1995 900m , 11°27′N 107°43′E Pacholatko & Dembicky”; NMW . – Son La Province • 11 ex. ; “N-Vietnam: Son La Prov., 36 km S Mac Chau [ Moc Chau ], ca. 1092 m , 20°44′34″N 104°55′44″E , 25.- 26.6.1997 , lg. C.L. [Chun-lin] LI”; NMW • 1 ex. ; “N-Vietnam: Son La Prov. , 36 km S Mac Chau [ Moc Chau ], ca. 1090 m , 20°45′34″N 104°55′44″E , 1.7.1997 , leg. C.F. [Chi-feng] Lee [it is not entirely clear, whether this specimen was collected in Son La Prov., because the coordinates lie in the Thanh Hóa Prov., very close to the boarder to Son La Prov., the label data (incl. the coordinates) are remarkably similar to the label data of the 11 specimens listed above, the elevation data seem to be incorrect in both cases]”; NMW . – Thanh Hóa Province • 36 ex. ; “ VIETNAM , Thanh Hóa Prov., Bá Thư ớc Distr. , Thành SƠn, Kho Mư ờng, 20°28′51.3″N , 105°07′51.6″E , 390 m a.s.l. , 11.11.2022 , Kodada & Selnekovič leg. (3)”; CKB . Type locality Thailand , Mae Hong Son Province , Ban Si Lang. Description Measurements (mm): TL: ♂♂ 2.90–3.33 (3.07± 0.12, n =26), ♀♀ 2.88–3.35 (3.20 ± 0.14, n =9); PL: ♂♂ 0.72–0.83 (0.75± 0.03, n= 26), ♀♀ 0.67–0.81 (0.76 ± 0.04, n=9); PW: ♂♂ 1.19–1.32 (1.25± 0.04, n= 26), ♀♀ 1.16–1.35 (1.29± 0.06, n =9); EL: ♂♂ 2.16–2.55 (2.32 ±0.10, n= 26), ♀♀ 2.21–2.55 (2.44 ± 0.11, n= 9); EW: ♂♂ 1.48–1.64 (1.56± 0.04, n=26), ♀♀ 1.48–1.69 (1.61 ± 0.06, n =9); PhL: 0.46–0.55 (0.50± 0.02, n =26); PrL: 0.28–0.35 (0.32 ± 0.31, n= 26). Body oblong-ovate, and dorsally convex ( Fig. 40 ). Integument black; mouthparts, antennae, trochanters, basal parts of femora, and tarsi reddish-brown; dorsum with fine bronze metallic lustre. Pubescence consisting of short yellowish setae. Dorsal plastron present on cranial surface (except for median part of vertex), anterior angles of pronotum, and on lateral sides of elytra. Ventral plastron absent on prosternal process, median part of metaventrite, and middle of ventrites 1–2. Dorsal surface of head with round setiferous punctures of diameters slightly smaller than an eye facet, separated by ca 0.5–2.0× puncture diameters. Plastron present on entire dorsal surface, except for oval area on vertex. Labrum transverse ( Fig. 1C ), anterior margin slightly emarginate in middle, exposed portion microreticulate, with small setiferous punctures; setae equal in length in both sexes. Anterior margin of clypeus straight. Eyes oval, moderately protruding from head outline, interfacetal setae short; ID: ♂♂ 0.39–0.46 mm (0.43 ± 0.01, n=26), ♀♀ 0.42–0.47 mm (0.44 ± 0.01, n=9). Antennae 10-segmented, densely setose. Pronotum ( Fig. 3E ) wider than long, widest at base, PW/PL: ♂♂ 1.59–1.76 (1.66± 0.03, n=26), ♀♀ 1.64–1.78 (1.70 ± 0.04, n=9); disc moderately convex; anterior angles protruding, acute; lateral sides convergent, slightly rounded in dorsal aspect; surface smooth, with round setiferous punctures; plastron confined to anterior angles. Prosternal process wider than long, lateral and distal edges rounded; lateral margins moderately raised, without clusters of long setae in males; median keel moderately arcuate. Metaventrite covered with plastron, except for slightly convex, posteriorly expanded median part; metaventral process with lateral margins raised, delimited by longitudinal grooves. Scutellum longer than wide, punctate. Elytra elongate oval, widest behind middle, EL/EW: ♂♂ 1.43–1.56 (1.49± 0.03, n= 26), ♀♀ 1.47–1.56 (1.51 ± 0.03, n =9); dorsal surface finely microreticulate; round setiferous punctures scattered on entire surface; plastron present on lateral portions forming a distally expanded band, anteriorly almost reaching elytral base, covering about one-third of elytron width (along elytral midlength). Tibiae slightly curved; protibia ca 1.2 × as long as protarsus; PrTL/PL: ♂♂ 1.08–1.22 (1.16 ± 0.06, n=26), ♀♀ 1.10–1.25 (1.16 ±0.05, n =9). Terminal protarsomere longer than the preceding segments combined. Abdomen covered with plastron, except on middle of first two ventrites. Apex of ventrite 5 rounded; females with small longitudinal keel before apex ( Fig. 4I ). Aedeagus ( Fig. 41 ): phallobase moderately long, slightly expanded proximally, PhL/PrL: 1.37–1.81 (1.57± 0.11, n=25); parameres weakly curved ventrad, apices narrowly rounded (lateral aspect); penis rounded apically; sclerotised fibula rather short and slender. Bursa copulatrix without microsclerites. Secondary sexual dimorphism Females possess a small longitudinal subapical keel on ventrite 5. Distribution China ( Guangdong , Hong Kong , Yunnan), Laos , Malaysia ( Perak ), Myanmar , Thailand , Vietnam ( Fig. 111A ).