An Appraisal of the Higher Classification of Cicadas (Hemiptera: Cicadoidea) with Special Reference to the Australian Fauna
Author
Moulds, MS
text
Records of the Australian Museum
2005
2005-11-30
57
3
375
446
https://doi.org/10.3853/j.0067-1975.57.2005.1447
journal article
263912
10.3853/j.0067-1975.57.2005.1447
0e5dc21c-0971-430f-95e2-8d3aa91ebeb0
2201-4349
8239161
Subfamily
Tettigadinae
Distant, 1905
Tibicinae
Buckton, 1889: 270
n.syn.
Tibicininae
sensu
Distant, 1905b: 304
(partim).
Tettigadinae
Distant, 1905d: 479
.
Tibicinidae
sensu
Metcalf, 1939: 247
(partim).
Platypediinae
Kato, 1932
n.syn.
If one follows the principle of priority (
Code
, Article 23) then the name
Tibicininae
should take priority as it is the oldest available family-group name associated with the genera included in this new subfamily concept
1
. However, the name
Tibicininae
previously represented a major family group of quite different concept as discussed at length throughout much of this paper. To now change its meaning would not only cause confusion in this regard but more importantly perpetuate a multitude of digressions and confusion previously associated with it and which are well documented by
Boulard (1988a
,
1998
,
2001
). I therefore take this opportunity to abandon the use of the name
Tibicininae
and its family-group derivatives and adopt the next available family-group name,
Tettigadinae
, a name which not only lacks a history of confusion but also retains its meaning to a large extent in this new concept. Official use of
Tettigadinae
requires suppression of the name
Tibicininae
and its derivatives by the International Commission of Zoology Nomenclature (
Code
, Article 19); a case for suppression is in preparation.
Type genus
.
Tettigades
Amyot & Serville
(type species:
T. chilensis
Amyot & Serville
).
Included tribes
. Platypediini Kato, Tettigadini Distant,
Tibicinini
Distant.
Diagnosis
. Metanotum entirely concealed at dorsal midline (except in
Platypedia
and
Tibicina
). Fore wing veins CuP and 1A unfused. Hind wing 1st cubital cell width at distal end about equal to 2nd cubital cell; veins RP and M unfused at their bases. Rim of abdominal timbal cavity with a turnedback rim. Pygofer with distal shoulder undeveloped; pygofer upper lobe absent. Uncus exceedingly long and nonretractable within pygofer. Claspers absent. Aedeagus with ventrobasal pocket present; aedeagal restrained by tubular encapsulation below uncus. Apical part of theca with a pair of leaf-like lateral lobes.
Distinguishing characters
. Fore wing veins CuP and 1A not fused. Hind wing veins RP and M unfused at base. Uncus exceedingly long and non-retractable within pygofer. Aedeagus with ventrobasal pocket present. Aedeagus with apical part of theca bearing a pair of leaf-like lateral lobes and a non-retractable tubular vesica. The male opercula have a distinctive S-shape, their lateral margins deeply concave, distal margins not reaching distal margins of tympanal cavities, basally not extending beyond meracantha.
Discussion
. This is an entirely new subfamily concept expanded to incorporate the former
Tettigadinae
,
Platypediinae
and the tribe
Tibicinini
. Previously identified attributes for the
Tettigadinae
in its former concept (
Distant, 1905d
;
Jacobi, 1907b
;
Chen, 1943
;
Boulard, 1976a
,
1986b
;
Hayashi, 1984
;
Duffels, 1993
;
Chou
et al
., 1997
) are now applicable at tribal rank for the Tettigadini.