Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)
Author
DeVries, Thomas J.
text
Journal of Natural History
2019
2018-12-17
53
25
1533
1584
journal article
10.1080/00222933.2018.1524032
034ebcd9-efa7-416a-822c-b6aee0e57a1c
1464-5262
3670229
Genus
Rhombopsis
Gardner, 1916
Type
species:
Fusus newberryi
Meek and Hayden, 1857
, by original designation. Cretaceous, Nebraska,
USA
.
Remarks
Rhombopsis
was proposed by
Gardner (
1916
, 456) in mid-paragraph, stating a belief that (1)
Maryland
‘
fulguroids
’
could not be placed in
Pyrifusus
Conrad, 1858
(s.s.); (2) that the
Maryland
taxa could be grouped with a subset of
Pyrifusus
species exhibiting sinuous outer lips and flattened columellas, as defined by
Meek (1876)
; (3) the subset deserved more than the subgeneric rank created by
Meek (1876)
; and (4) Meek
’
s (1876) name for the species subset,
Neptunella
subgen. nov.
, was preoccupied by
Neptunella
Gray,
1854,
a ranellid gastropod, and so needed to be replaced
–
by
Rhombopsis
. The reader might infer that subsequently described fulguroid
Maryland
species (
Gardner
1916
, 457
–
463) were considered examples of the newly elevated and renamed
Neptunella
, i.e.
Rhombopsis
, although
Gardner (1916)
still used
Pyrifusus
in her description of the
Maryland
species:
P. marylandicus
sp. nov.
,
P. vittatus
sp. nov.
,
P. monmouthensis
sp. nov.
,
P. cuneus
Whitfield,
1892,
P. whitfieldi
sp. nov.
, and
P. elevata
Whitfield, 1892
.
Sohl (1964)
addressed the scope of the genus
Rhombopsis
in his monograph on Late Cretaceous molluscs from
Tennessee
and
Mississippi
. From a number of
Maryland
species,
Sohl (
1964
, 196) accepted only
Pyrifusus marylandicus
Gardner, 1916
as an example of
Rhombopsis
. Later on,
Sohl (
1964
, 199) distinguished between
Rhombopsis
and the closely related
Deussenia
Stephenson, 1941
, on the basis of the presence of a strongly developed subsutural collar (weak or absent on the former genus, strong on the latter genus). Nevertheless,
Sohl (1964)
proceeded to describe a new species,
R. molinoensis
, with a broad subsutural collar. The figures of
R. molinoensis
(
Sohl
1964
, pl. 24, figs. 14, 15) are biconic, rather than fusiform, and have a uniformity of spiral sculpture (narrow primary spiral cords from the base of the subsutural collar to the tip of the siphonal canal, all crossing the axial ribs) also seen in the
type
species of
Rhombopsis
,
R. newberryi
(see
Stanton
1920
, pl. 7, fig. 8a, 8b). These characters are not seen in a figure of the Late Cretaceous species from Tennessee,
Rhombopsis
?
[sic]
orientalis
Wade, 1926
(
Sohl
1964
, pl. 24, fig. 5), nor in figure of the Late Cretaceous
R. gracilis
(Stanton, 1920)
from
North Dakota
.
Sohl (
1964
, 199) questioned Olsson
’
s (1944) assignment to
Rhombopsis
of the Late Cretaceous species,
R. meridionalis
Olsson, 1944
, from the
Tortuga
Formation of northern Peru, based on a siphonal canal curved to a greater extent than seen in North American Cretaceous species.
The proper separation of these small Late Cretaceous fasciolariids (or melongenids) is beyond the scope of this study. It can be noted that among the taxa falling within the scope of discussion addressing
Rhombopsis
, there is a subset of species with fusiform profiles, a weak or absent subsutural collar, and primary and secondary spiral cords:
R. gracilis
,
R. orientalis
(Tennessee),
R. marylandicus
(Maryland), and
R. meridionalis
(northern
Peru
). Late Cretaceous specimens from the central Peruvian Andes tentatively assigned to
Rhombopsis
by
Berry and Singwald (1922)
do not resemble any taxa of
Rhombopsis
, whether the genus is defined broadly or narrowly.