Records about the alien millipede Oxidus gracilis (C. L. Koch, 1847) (Diplopoda: Polydesmida: Paradoxosomatidae) in continental Chile
Author
Parra-Gómez, Antonio
Facultad de Ciencias, Universidad Austral de Chile, Av. Rector Eduardo Morales Miranda 23, Valdivia, Región de Los Ríos.
pg.antonio@hotmail.com
text
Revista Chilena de Entomología
2022
Rev. Chil. Entomol.
2022-01-31
48
1
73
79
http://dx.doi.org/10.35249/rche.48.1.22.06
journal article
294445
10.35249/rche.48.1.22.06
9a9baa51-df77-492a-91db-60784836f175
0718-8994
10950238
9FE1BC5D-6228-48F9-93CF-E867E041709F
Oxidus gracilis
(C.L.
Koch, 1847
)
(
Figs. 1-4
)
Fontaria gracilis
C.L.
Koch 1847: 142
.
Oxidus gracilis
:
Verhoeff 1924: 416
, fig. 18;
Chamberlin 1957: 12
.
Orthomorpha gracilis
:
Skottsberg 1956: 418
.
Material examined.
Oxidus gracilis
.
1 male
.
CHILE
.
Coquimbo
,
Choapa
,
Salamanca
, in a water fountain
MLP
(sic.),
17-V-2011
, col.
P.
Baéz, B
. Collado and
S. Letieler
[
MHNH
].
5 females,
5 males
. Limarí,
Bosque Fray Jorge National Park
, under a log close to reedbed, park administration,
30°40’31.12” S
-
71°38’9.57” W
,
5-X-2017
, col.
Jorge Pérez Schultheiss
[
MHNH
].
9 females
,
5 males
.
Santiago
, Linderos, Buin, under ornamental stones in a house garden,
33°45’09” S
-
70°44’16” W
,
16-I-2019
, e-13 (field code), col.
Patricio Farah
[
APG
]
.
Additional records:
CHILE
.>
10 specimens
(sic.).
Valparaíso
,
Quillota
,
La Cruz
,
18-I-1973
,
Col. Walter C. Sedgwick.
Det. Herbet W. Levi
(presumably) (Coordinates incorrectly given as “
34°30’30.6720” S
-
71°53’45.7404” W
”) (
MCZBASE 2021
)
.
Figures 1-2.
Oxidus gracilis
. 1. Lateral view, female. 2. Dorsal view, male,
collum
broken. Scale: 1 mm. /
Oxidus gracilis
.
1. Vista lateral, hembra. 2. Vista dorsal, macho,
collum
roto. Escala: 1 mm.
Descriptive notes.
Length ca
18-19 mm
in males, ca
20-21 in
females. Width of midbody pro- and metazonae in males and females ca 1.3-1.4 and ca
1.9-2 mm
respectively. General coloration dark brown with yellowish paranota; legs also yellowish. Body with 20 segments. Antennae long, extending back behind 2nd segment when stretched dorsally (a little shorter on females). Terga with transversal sulcus. Paraterga developed, anterolateral margin rounded, posterolateral margin dentiform. Epiproct tip bilobulated. Gonopods as
Nguyen
et al.
(2017)
and
Rojas-Buffet
et al.
(2020)
: Femorite (fe) expanded distally; postfemoral lamina (l) with a rectangular shape; postfemoral spine (z) tubercular and spiniform; postfemoral process (h) thin and serrated at the distolateral part; basal part of process (h) and spine (z) noticeably separated; solenophore (sph) with a lamelliform mesal lobule; tips of mesal lobule and solenophore circular (
Fig. 3
).
Remarks.
The species differs from
Ologonosoma
Silvestri, 1897
congeners, by the presence of prominent paranota with yellowish lateral margin (
Figs. 1-2
).
Oxidus gracilis
is widely distributed over the world primarily in warm temperate areas (
Nguyen
et al
. 2017
). The material from
Chile
has been collected in Rapa Nui (Easter Island) (
Verhoeff 1924
) and also in continental
Chile
in 1973 by Walter C. Sedgwick, as other invertebrates were collected in the same date and locality by this author and possibly identified by Herbet W. Levi as he worked with spiders from the same batch (
MCZBASE 2021
,
2022
), no subsequent publication has been issue citing this last sample. Although records from
GBIF
show
O. gracilis
to be a very common species in central
Chile
(
GBIF
2021), especially in urban environments, it is worth noting that some of our specimens were collected in the park administration zone of the Bosque Fray Jorge National Park, near manmade structures (
Fig. 4
).
Apart from
Oxidus
, only one
Paradoxosomatidae
genus from
Chile
is known:
Ologonosoma
(
Fig. 5
). Species from this genus have been recorded from Bosque Fray Jorge National Park and
Valparaíso
in the case of
Ologonosoma robustum
(
Attems, 1898
)
and Valdivia and nearby places for
Ologonosoma ecarinatum
(
Attems, 1898
)
and
Ologonosoma primum
(
Chamberlin, 1957
)
, all 3 of them endemic to the Chilean territory (
Attems 1898
,
1903
;
Chamberlin 1957
;
Silva & Sáiz 1975
; Parra-Gómez unpublished). Only
O. rubustum
seems to overlap its known distribution with the records of
O. gracilis
. Impacts of exotic diplopods on native species in
Chile
is still unknown as no study has ever been conducted in the country.
Figure 3.
Male right gonopod view under compound microscope, lateral view. Nomenclature by
Nguyen
et al.
(2017)
: h = postfemoral process; sph = solenophore; sl = solenomere; z = postfemoral spine; l = postfemoral lamina; fe = femorite; co = coxa; pf = prefemorite. Scale: 0.1 mm / Gonópodo derecho del macho visto bajo microscopio compuesto, vista lateral. Nomenclatura por
Nguyen
et al.
(2017)
: h = proceso postfemoral; sph = solenóforo; sl = solenómero; z = espina postfemoral; l = lámina postfemoral; fe = femorito; co = coxa; pf = prefemorito. (nota: la translación de algunos términos al español es tentativa, debido a que en muchos casos no existe una traducción oficial). Escala: 0,1 mm.
According to the records known in the literature, the exotic milliped fauna of continental
Chile
contains a total of six species:
B. guttulatus
(Bosc, 1792)
,
B. pusillus
(Leach, 1814)
,
C. britannicus
(Verhoeff, 1891)
,
C. latestriatus
(Curtis, 1845)
,
N. kochii
(Gervais, 1847)
(
Attems 1903
;
Verhoeff 1924
;
Schubart 1964
), and
O. gracilis
which extends from Caballo Muerto near Ovalle to Chillán, based on the additional records herein (
Fig. 4
).