A new genus Gnathospinosa (Lepidoptera: Tineidae: Euplocaminae) from China, with description of a new species and its taxonomic position Author Liao, Cheng-Qing Author Wang, Min Author Huang, Guo-Hua text Zootaxa 2019 2019-02-14 4555 3 416 424 journal article 27540 10.11646/zootaxa.4555.3.10 ea697764-3953-4089-a7f5-e392e60dd850 1175-5326 2624396 550598E8-4E22-44F3-AC5E-F68A44638E6A Gnathospinosa Liao & Huang , gen. nov. Type species: Gnathospinosa qinlingensis Liao & Huang , sp. nov. by present designation. Description. Male. Wing span about 17–18 mm . Head : Vertex with erect piliform scales. Antenna bipectinate, pectinations elongate and ciliated; scape elongate, cylindrical; pedicel short; flagellum with 35 flagellomeres, basal 25 flagellomeres bipectinate. Maxillary palpus with 3 segments, very short and indistinct. Galeae short. Labial palpus developed and directed forward, second palpomere expanded, lateral bristles absent. Thorax : Dorsal surface covered with yellowish scales mixed some black scales. Foreleg with epiphysis present on tibiae ( Fig. 4D ), tarsus without spines; occurrence of spurs on legs follows the formula 0-2-4 ( Figs. 4 A–C). Wings moderately elongate, slightly rounded apically; forewings ground color yellowish with large dark spots; venation complete, R 4 and R 5 long stalked in forewing; R 5 ending at termen near apex; chorda present; 1A+2A separated at base; hindwing relatively elongate, rounded apically, all veins present; crossvein M-CuA present, M branched in discal cell ( Fig. 2 ). Abdomen : Male genitalia complex; uncus simple, divided into two lobes; tegumen and vinculum fused into a ring; gnathos broad, strongly sclerotized, covered with spines; saccus absent; aedeagus small, inflated basally, without cornuti. Female unknown. Distribution. China ( Shaanxi ). Etymology. The generic name is derived from the structure of male genitalia with the gnathos covered with scaly spines ( spinosus in Latin). The generic name should be treated as feminine. Remarks. This genus is placed in the subfamily Euplocaminae according to the following morphological evidences: in males i) antenna bipectinate; ii) valva two-folded, iii) uncus bilobed, iv) gnathos arms fused medially ( Gaedike 2015 ). It can easily be distinguished from other subfamilies in Tineidae by the following diagnostic characters in males: i) valva with an auricular hairy projection on inner surface, ii) gnathos broad, strongly sclerotized and covered with scaly spines.