<document id="EBF39401B021EB10128081AAD8A0C04C" ID-DOI="10.11646/zootaxa.4809.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3933989" ID-ZooBank="6E4829CF-1476-4FDB-941D-9EAFCA29D011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594125923415" checkinUser="plazi" docAuthor="Fend, Steven &amp; Rodriguez, Pilar" docDate="2020" docId="03D287B7A572FFC2FF7D6A15F14EFCA2" docLanguage="en" docName="zootaxa.4809.1.6.pdf" docOrigin="Zootaxa 4809 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eremidrilus gilita Fend &amp; Rodriguez 2020, n. sp." docType="treatment" docUuid="C090F3C9-BBF0-4FCF-A8BD-517C92FDC4F5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="124" masterDocId="FFEBFFCFA578FFCFFFEA6D45F50DFFB5" masterDocTitle="New Eremidrilus species (Clitellata: Lumbriculidae) from western North America Part 1, species with two spermathecal segments" masterLastPageNumber="131" masterPageNumber="111" pageNumber="121" updateTime="1698778818868" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="05A1EB7A8DF46382CF9CEF048DAB3275">New Eremidrilus species (Clitellata: Lumbriculidae) from western North America Part 1, species with two spermathecal segments</mods:title>
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<mods:namePart id="912DFF56E26E91F9F0ED6FDB69153ADA">Fend, Steven</mods:namePart>
<mods:affiliation id="3F6A018057579E791D1CC6A83F60BAC0">85060 Spencer Hollow Rd., Eugene, OR, 97405, USA.</mods:affiliation>
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<mods:namePart id="92A309254770DC013DB73202DFCBD33C">Rodriguez, Pilar</mods:namePart>
<mods:affiliation id="8378D17E6E3E04AD6E06D411AC07C347">Zoology and Animal Cell Biology Dpt., Faculty of Science and Technology, University of the Basque Country UPV / EHU, Apdo. 644, Bilbao 48080, Spain. pilar. rodriguez @ ehu. es; https: // orcid. org / 0000 - 0001 - 8387 - 1669</mods:affiliation>
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<mods:date id="422AE030501A86A4EDD40357CF46A7A1">2020</mods:date>
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<paragraph id="8BC436A1A572FFC5FF7D6A15F4BAF8DE" blockId="10.[151,439,1872,1935]" box="[151,439,1872,1899]" pageId="10" pageNumber="121">
<heading id="D08C81CDA572FFC5FF7D6A15F4BAF8DE" bold="true" box="[151,439,1872,1899]" fontSize="11" level="1" pageId="10" pageNumber="121" reason="1">
<emphasis id="B90FEAB3A572FFC5FF7D6A15F4BAF8DE" bold="true" box="[151,439,1872,1899]" pageId="10" pageNumber="121">
<taxonomicName id="4C7B4D22A572FFC5FF7D6A15F463F8DE" ID-CoL="869VY" authority="Fend &amp; Rodriguez, 2020" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[151,366,1872,1899]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="10" pageNumber="121" phylum="Annelida" rank="species" species="gilita" status="sp. nov.">
<emphasis id="B90FEAB3A572FFC5FF7D6A15F463F8DE" bold="true" box="[151,366,1872,1899]" italics="true" pageId="10" pageNumber="121">Eremidrilus gilita</emphasis>
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<taxonomicNameLabel id="A23C57C8A572FFC5FE9F6A14F4BAF8DE" box="[373,439,1873,1899]" pageId="10" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C361652AA572FFC5FF7D6A30F465F83A" box="[151,360,1909,1935]" pageId="10" pageNumber="121" type="description">
<paragraph id="8BC436A1A572FFC5FF7D6A30F465F83A" blockId="10.[151,439,1872,1935]" box="[151,360,1909,1935]" pageId="10" pageNumber="121">
(
<figureCitation id="13402A24A572FFC5FF756A30F401F83A" box="[159,268,1909,1935]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="10" pageNumber="121">Figures 7</figureCitation>
, 
<figureCitation id="13402A24A572FFC5FEF26A30F42AF83A" box="[280,295,1909,1935]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="10" pageNumber="121">8</figureCitation>
, 
<figureCitation id="13402A24A572FFC5FEDE6A30F46DF83A" box="[308,352,1909,1935]" captionStart="FIGURE 11" captionStartId="18.[151,250,1571,1596]" captionTargetBox="[281,1342,238,1481]" captionTargetId="figure@18.[200,1387,199,1548]" captionTargetPageId="18" captionText="FIGURE 11. Eremidrilus species having two spermathecal segments, schematic illustration comparing position and basic morphology of reproductive organs in X–XIII." figureDoi="http://doi.org/10.5281/zenodo.3934011" httpUri="https://zenodo.org/record/3934011/files/figure.png" pageId="10" pageNumber="121">11E</figureCitation>
)
</paragraph>
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<paragraph id="8BC436A1A572FFC5FF7D6AF8F7CBF84E" blockId="10.[151,1436,1981,2043]" pageId="10" pageNumber="121">
<materialsCitation id="3B133CFCA572FFC5FF7D6AF8F7CBF84E" collectionCode="USNM" country="Canada" location="Dissected" pageId="10" pageNumber="121" specimenCode="USNM 1618768" specimenCount="1" typeStatus="holotype">
<emphasis id="B90FEAB3A572FFC5FF7D6AF8F406F862" bold="true" box="[151,267,1981,2007]" pageId="10" pageNumber="121">
<typeStatus id="54C08803A572FFC5FF7D6AF8F40BF862" box="[151,262,1981,2007]" pageId="10" pageNumber="121" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBDD9EDAA572FFC5FEFE6AF8F4DEF862" box="[276,467,1981,2007]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="121" type="Museum">USNM 1618768</specimenCode>
. 
<location id="8EA4607AA572FFC5FE356AF8F741F862" LSID="urn:lsid:plazi:treatment:03D287B7A572FFC2FF7D6A15F14EFCA2:8EA4607AA572FFC5FE356AF8F741F862" box="[479,588,1981,2007]" country="Canada" name="Dissected" pageId="10" pageNumber="121">Dissected</location>
worm, incomplete but with sperm in spermathecae and mature egg; slidemounted in 
<collectingCountry id="F36C7631A572FFC5FEF76AA4F47CF84E" box="[285,369,2017,2043]" name="Canada" pageId="10" pageNumber="121">Canada</collectingCountry>
balsam, stained with carmine.
</materialsCitation>
</paragraph>
<paragraph id="8BC436A1A573FFC4FF2D6DDCF4CCFF62" blockId="11.[151,1436,152,359]" pageId="11" pageNumber="122">
<emphasis id="B90FEAB3A573FFC4FF2D6DDCF47CFF06" bold="true" box="[199,369,152,179]" pageId="11" pageNumber="122">Type Locality.</emphasis>
<materialsCitation id="3B133CFCA573FFC4FE926DDCF4CCFF62" collectingDate="1996-04-22" collectorName="S. Fend." country="United States of America" county="Catron Co." location="Willow Creek" municipality="Gilita Creek" pageId="11" pageNumber="122" specimenCount="1" stateProvince="New Mexico">
<collectingCountry id="F36C7631A573FFC4FE926DDCF4BEFF06" box="[376,435,153,179]" name="United States of America" pageId="11" pageNumber="122">USA</collectingCountry>
, 
<collectingRegion id="49BFF843A573FFC4FE546DDCF742FF06" box="[446,591,153,179]" country="United States of America" name="New Mexico" pageId="11" pageNumber="122">New Mexico</collectingRegion>
, 
<collectingCounty id="62A54E2DA573FFC4FDB06DDCF7D9FF06" box="[602,724,153,179]" pageId="11" pageNumber="122">Catron Co.</collectingCounty>
, 
<collectingMunicipality id="6BA0ACDBA573FFC4FD356DDCF669FF06" box="[735,868,153,179]" pageId="11" pageNumber="122">Gilita Creek</collectingMunicipality>
at 
<location id="8EA4607AA573FFC4FC6F6DDCF113FF06" LSID="urn:lsid:plazi:treatment:03D287B7A572FFC2FF7D6A15F14EFCA2:8EA4607AA573FFC4FC6F6DDCF113FF06" box="[901,1054,153,179]" country="United States of America" county="Catron Co." municipality="Gilita Creek" name="Willow Creek" pageId="11" pageNumber="122" stateProvince="New Mexico">Willow Creek</location>
(Site 14, 
<tableCitation id="C6F9031AA573FFC4FB6D6DDCF1DBFF06" box="[1159,1238,153,179]" captionStart="TABLE 1" captionStartId="15.[151,239,668,694]" captionText="TABLE 1. Eremidrilus n. spp., collection sites." pageId="11" pageNumber="122">Table 1</tableCitation>
), in riffle, 
<date id="FFC51061A573FFC4FABB6DDCF5DDFF62" pageId="11" pageNumber="122" value="1996-04-22">
<collectingDate id="EF81E989A573FFC4FABB6DDCF5DDFF62" pageId="11" pageNumber="122" value="1996-04-22">22 Apr 1996</collectingDate>
</date>
, collected by 
<collectorName id="268E5377A573FFC4FE8C6DF8F4CCFF62" box="[358,449,189,215]" pageId="11" pageNumber="122">S. Fend.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BC436A1A573FFC4FF2D6DA4F6ECFEF6" blockId="11.[151,1436,152,359]" pageId="11" pageNumber="122">
<materialsCitation id="3B133CFCA573FFC4FF2D6DA4F6D0FEF6" collectionCode="USNM, MNCN" country="Canada" location="All" pageId="11" pageNumber="122" specimenCode="USNM 1618770, 1, USNM 1618769, MNCN 16.3" specimenCount="1" typeStatus="paratype">
<emphasis id="B90FEAB3A573FFC4FF2D6DA4F44BFF4E" bold="true" box="[199,326,225,251]" pageId="11" pageNumber="122">
<typeStatus id="54C08803A573FFC4FF2D6DA4F44CFF4E" box="[199,321,225,251]" pageId="11" pageNumber="122" type="paratype">Paratypes</typeStatus>
.
</emphasis>
From the type locality (Site 14), same collection data; all mature: 
<specimenCode id="DBDD9EDAA573FFC4FBBA6DA4F01DFF4E" box="[1104,1296,225,251]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="122" type="Museum">USNM 1618770</specimenCode>
, 
<specimenCode id="DBDD9EDAA573FFC4FAF46DA4F021FF4E" box="[1310,1324,225,251]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="122" type="Museum">1</specimenCode>
sagittally sectioned worm on 2 slides, stained with hematoxylin and eosin; 
<specimenCode id="DBDD9EDAA573FFC4FC846C40F126FEAA" box="[878,1067,261,287]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="122" type="Museum">USNM 1618769</specimenCode>
, 
<specimenCode id="DBDD9EDAA573FFC4FBDF6C40F14EFEAA" box="[1077,1091,261,287]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="122" type="Museum">1</specimenCode>
dissected, incomplete. 
<specimenCode id="DBDD9EDAA573FFC4FAA26C40F5CAFEF6" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="122" type="Museum">MNCN 16.3</specimenCode>
/3115, 1 whole-mount, complete. 
<location id="8EA4607AA573FFC4FDAA6C6CF769FEF6" LSID="urn:lsid:plazi:treatment:03D287B7A572FFC2FF7D6A15F14EFCA2:8EA4607AA573FFC4FDAA6C6CF769FEF6" box="[576,612,297,323]" country="Canada" name="All" pageId="11" pageNumber="122">All</location>
slide-mounted in 
<collectingCountry id="F36C7631A573FFC4FCC56C6CF68EFEF6" box="[815,899,297,323]" name="Canada" pageId="11" pageNumber="122">Canada</collectingCountry>
balsam
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361652AA573FFC4FF2D6C08F106FED2" box="[199,1035,333,359]" pageId="11" pageNumber="122" type="etymology">
<paragraph id="8BC436A1A573FFC4FF2D6C08F106FED2" blockId="11.[151,1436,152,359]" box="[199,1035,333,359]" pageId="11" pageNumber="122">
<emphasis id="B90FEAB3A573FFC4FF2D6C08F443FED2" bold="true" box="[199,334,333,359]" pageId="11" pageNumber="122">Etymology.</emphasis>
Named for the 
<typeStatus id="54C08803A573FFC4FE176C08F720FED2" box="[509,557,333,359]" pageId="11" pageNumber="122">type</typeStatus>
locality, Gilita Creek. Noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF046629A573FFC4FF7D6BF8F6ADF8AB" ID-DOI="http://doi.org/10.5281/zenodo.3934003" ID-Zenodo-Dep="3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="11" pageNumber="122" startId="11.[151,250,1725,1750]" targetBox="[162,1421,391,1687]" targetPageId="11">
<paragraph id="8BC436A1A573FFC4FF7D6BF8F6ADF8AB" blockId="11.[151,1437,1725,1822]" pageId="11" pageNumber="122">
<emphasis id="B90FEAB3A573FFC4FF7D6BF8F41EF963" bold="true" box="[151,275,1725,1750]" pageId="11" pageNumber="122">FIGURE 7.</emphasis>
<taxonomicName id="4C7B4D22A573FFC4FEF36BFBF4C0F963" authority="Fend &amp; Rodriguez, 2020" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[281,461,1726,1750]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="11" pageNumber="122" phylum="Annelida" rank="species" species="gilita" status="sp. nov.">
<emphasis id="B90FEAB3A573FFC4FEF36BFBF4C0F963" box="[281,461,1726,1750]" italics="true" pageId="11" pageNumber="122">Eremidrilus gilita</emphasis>
</taxonomicName>
<emphasis id="B90FEAB3A573FFC4FE396BFBF701F963" bold="true" box="[467,524,1726,1750]" pageId="11" pageNumber="122">
<taxonomicNameLabel id="A23C57C8A573FFC4FE396BFBF701F963" box="[467,524,1726,1750]" pageId="11" pageNumber="122" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, from Gilita Creek. 
<emphasis id="B90FEAB3A573FFC4FD336BF8F7FCF963" bold="true" box="[729,753,1725,1750]" pageId="11" pageNumber="122">A.</emphasis>
Anterior ends, external view of unmounted worms, both with pharynx partially everted. 
<emphasis id="B90FEAB3A573FFC4FE9E6BA7F486F94F" bold="true" box="[372,395,1762,1786]" pageId="11" pageNumber="122">B.</emphasis>
Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. 
<emphasis id="B90FEAB3A573FFC4FC066BA4F109F94F" bold="true" box="[1004,1028,1761,1786]" pageId="11" pageNumber="122">C.</emphasis>
Reproductive organs, segments X–XIII. 
<emphasis id="B90FEAB3A573FFC4FF7D6A43F5BDF8AB" bold="true" box="[151,176,1798,1822]" pageId="11" pageNumber="122">D.</emphasis>
Reproductive organs, segments IX–XI (second spermatheca was broken).
</paragraph>
</caption>
<subSubSection id="C361652AA573FFC2FF2D6A0EF07EFDAE" lastPageId="13" lastPageNumber="124" pageId="11" pageNumber="122" type="description">
<paragraph id="8BC436A1A573FFC3FF2D6A0EF1C5FF62" blockId="11.[151,1436,1867,2039]" lastBlockId="12.[151,1437,153,359]" lastPageId="12" lastPageNumber="123" pageId="11" pageNumber="122">
<emphasis id="B90FEAB3A573FFC4FF2D6A0EF454F8D3" bold="true" box="[199,345,1867,1894]" pageId="11" pageNumber="122">Description.</emphasis>
Length of one complete, preserved worm 
<quantity id="4C839B44A573FFC4FCA86A09F698F8D2" box="[834,917,1868,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="122" unit="mm" value="17.0">17 mm</quantity>
, 58 segments. Diameter at clitellum 
<quantity id="4C839B44A573FFC4FAD06A0EF5CBF83E" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="11" pageNumber="122" unit="mm" value="0.425" valueMax="0.5" valueMin="0.35">0.35–0.5 mm</quantity>
. Proboscis 
<quantity id="4C839B44A573FFC4FEAD6A35F4DAF83E" box="[327,471,1904,1931]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" metricValueMax="4.8" metricValueMin="2.6" pageId="11" pageNumber="122" unit="mm" value="370.0" valueMax="480.0" valueMin="260.0">260–480 µm</quantity>
long, about 
<quantity id="4C839B44A573FFC4FD8C6A2AF7D6F83E" box="[614,731,1903,1931]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="11" pageNumber="122" unit="mm" value="90.0" valueMax="95.0" valueMin="85.0">85–95 µm</quantity>
diameter at middle (
<figureCitation id="13402A24A573FFC4FC2D6A35F12FF83F" box="[967,1058,1904,1930]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="11" pageNumber="122">Fig. 7A</figureCitation>
, 
<figureCitation id="13402A24A573FFC4FBC46A35F159F83F" box="[1070,1108,1904,1930]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="11" pageNumber="122">8A</figureCitation>
). Secondary annulation from IV to clitellum (
<figureCitation id="13402A24A573FFC4FEA06AD1F4A9F81B" box="[330,420,1940,1966]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="11" pageNumber="122">Fig. 7A</figureCitation>
). Chaetae with nodulus 0.4 to 0.5 from distal end in mid-body, but may be more distal posteriorly; length 
<quantity id="4C839B44A573FFC4FEAD6AFDF4D5F866" box="[327,472,1976,2003]" metricMagnitude="-1" metricUnit="m" metricValue="1.225" metricValueMax="1.33" metricValueMin="1.1199999999999999" pageId="11" pageNumber="122" unit="mm" value="122.5" valueMax="133.0" valueMin="112.0">112–133 µm</quantity>
, shorter in II (
<figureCitation id="13402A24A573FFC4FD916AFDF7D3F866" box="[635,734,1976,2003]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="11" pageNumber="122">Figs. 7B</figureCitation>
, 
<figureCitation id="13402A24A573FFC4FD006AFDF603F867" box="[746,782,1976,2002]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="11" pageNumber="122">8B</figureCitation>
). Male pores on the ventral chaetal line, midway between chaetae and posterior septum, on narrow porophores (
<figureCitation id="13402A24A573FFC4FD176A99F671F843" box="[765,892,2012,2038]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="11" pageNumber="122">Figs. 7A,C</figureCitation>
, 
<figureCitation id="13402A24A573FFC4FC636A99F6C4F843" box="[905,969,2012,2038]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="11" pageNumber="122">8C,D</figureCitation>
). Simple spermathecal pores (
<figureCitation id="13402A24A573FFC4FACA6A99F083F843" box="[1312,1422,2012,2038]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="11" pageNumber="122">Fig. 8E,F</figureCitation>
), behind ventral chaetae, on the ventral chaetal line, the first at 1/3 the distance from posterior septum to ventral chaetae, second close to intersegment 12/13 (
<figureCitation id="13402A24A574FFC3FDB56DF8F7CDFF62" box="[607,704,189,215]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="12" pageNumber="123">Figs. 7C</figureCitation>
, 
<figureCitation id="13402A24A574FFC3FD276DF8F606FF62" box="[717,779,189,215]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="12" pageNumber="123">8E,H</figureCitation>
). Female pores intersegmental at 11/12.
</paragraph>
<paragraph id="8BC436A1A574FFC3FF2D6DA4F496FED2" blockId="12.[151,1437,153,359]" pageId="12" pageNumber="123">
Epidermis 
<quantity id="4C839B44A574FFC3FEAA6DA5F4A9FF4E" box="[320,420,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="12" pageNumber="123" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 µm</quantity>
thick, up to 
<quantity id="4C839B44A574FFC3FDC66DA4F779FF4E" box="[556,628,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="123" unit="mm" value="12.0">12 µm</quantity>
in clitellum. Clitellum X to XIII. Pharynx in II–IV, dorsal pad not prominent; pharyngeal glands in IV–VI (VII). Sperm sacs extend anteriorly to VII, posteriorly to XIII or XIV. Egg sacs to XV. Nephridia in several postclitellar segments, terminating in a round vesicle (
<quantity id="4C839B44A574FFC3FBD96C6CF173FEF6" box="[1075,1150,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="123" unit="mm" value="30.0">30 µm</quantity>
wide), anterior to ventral chaetae (
<figureCitation id="13402A24A574FFC3FF106C08F451FED2" box="[250,348,333,359]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="12" pageNumber="123">Figs. 7C</figureCitation>
, 
<figureCitation id="13402A24A574FFC3FE826C08F483FED2" box="[360,398,333,359]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="12" pageNumber="123">8G</figureCitation>
).
</paragraph>
<caption id="DF046629A574FFC3FF7D6A5FF7A5F876" ID-DOI="http://doi.org/10.5281/zenodo.3934005" ID-Zenodo-Dep="3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="12" pageNumber="123" startId="12.[151,250,1818,1843]" targetBox="[151,1433,381,1792]" targetPageId="12">
<paragraph id="8BC436A1A574FFC3FF7D6A5FF7A5F876" blockId="12.[151,1436,1818,1987]" pageId="12" pageNumber="123">
<emphasis id="B90FEAB3A574FFC3FF7D6A5FF41EF886" bold="true" box="[151,275,1818,1843]" pageId="12" pageNumber="123">FIGURE 8.</emphasis>
<taxonomicName id="4C7B4D22A574FFC3FEF36A5EF4C3F886" authority="Fend &amp; Rodriguez, 2020" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[281,462,1819,1843]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="12" pageNumber="123" phylum="Annelida" rank="species" species="gilita" status="sp. nov.">
<emphasis id="B90FEAB3A574FFC3FEF36A5EF4C3F886" box="[281,462,1819,1843]" italics="true" pageId="12" pageNumber="123">Eremidrilus gilita</emphasis>
</taxonomicName>
<emphasis id="B90FEAB3A574FFC3FE3E6A5EF700F886" bold="true" box="[468,525,1819,1843]" pageId="12" pageNumber="123">
<taxonomicNameLabel id="A23C57C8A574FFC3FE3E6A5EF700F886" box="[468,525,1819,1843]" pageId="12" pageNumber="123" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, from Gilita Creek. 
<emphasis id="B90FEAB3A574FFC3FD306A5FF7FEF886" bold="true" box="[730,755,1818,1843]" pageId="12" pageNumber="123">A.</emphasis>
Anterior end of a whole-mounted worm. 
<emphasis id="B90FEAB3A574FFC3FB736A5EF1BDF886" bold="true" box="[1177,1200,1819,1843]" pageId="12" pageNumber="123">B.</emphasis>
Anterior dorsal chaeta, from a whole-mount. 
<emphasis id="B90FEAB3A574FFC3FE9D6A7BF482F8E2" bold="true" box="[375,399,1854,1879]" pageId="12" pageNumber="123">C.</emphasis>
Male porophore with ectal end of atrial duct, from a whole-mount. 
<emphasis id="B90FEAB3A574FFC3FBA26A7AF16CF8E2" bold="true" box="[1096,1121,1855,1879]" pageId="12" pageNumber="123">D.</emphasis>
Atrium and porophore, from a sagittal section. 
<emphasis id="B90FEAB3A574FFC3FED56A26F45BF8CE" bold="true" box="[319,342,1891,1915]" pageId="12" pageNumber="123">E.</emphasis>
Spermathecae from a sagittal section. 
<emphasis id="B90FEAB3A574FFC3FD036A26F7F1F8CE" bold="true" box="[745,764,1891,1915]" pageId="12" pageNumber="123">F.</emphasis>
Spermathecal duct, sagittal section. 
<emphasis id="B90FEAB3A574FFC3FB936A27F19EF8CE" bold="true" box="[1145,1171,1890,1915]" pageId="12" pageNumber="123">G.</emphasis>
Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. 
<emphasis id="B90FEAB3A574FFC3FD536AC2F7DEF82A" bold="true" box="[697,723,1927,1951]" pageId="12" pageNumber="123">H.</emphasis>
Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left.
</paragraph>
</caption>
<paragraph id="8BC436A1A575FFC2FF2D6DDCF103FE3E" blockId="13.[151,1437,152,791]" pageId="13" pageNumber="124">
Atria 
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long, entirely in X, club-shaped, maximum diameter 
<quantity id="4C839B44A575FFC2FC046DDDF169FF06" box="[1006,1124,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" metricValueMax="7.7" metricValueMin="5.2" pageId="13" pageNumber="124" unit="mm" value="64.5" valueMax="77.0" valueMin="52.0">52–77 µm</quantity>
; muscle layer thin, 
<quantity id="4C839B44A575FFC2FAD46DDCF095FF06" box="[1342,1432,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="13" pageNumber="124" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 µm</quantity>
; epithelial cells cuboidal, to 
<quantity id="4C839B44A575FFC2FE396DF8F721FF62" box="[467,556,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="13" pageNumber="124" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 µm</quantity>
high. Atrial ampulla covered with a prostate layer, 
<quantity id="4C839B44A575FFC2FB946DF8F1FEFF62" box="[1150,1267,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="124" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 µm</quantity>
thick, prostate cells in densely packed clusters (
<figureCitation id="13402A24A575FFC2FDED6DA4F78BFF4E" box="[519,646,225,251]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="13" pageNumber="124">Figs. 7C,D</figureCitation>
, 
<figureCitation id="13402A24A575FFC2FD7B6DA4F7BAFF4E" box="[657,695,225,251]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="13" pageNumber="124">8D</figureCitation>
). Atrium narrows to 
<quantity id="4C839B44A575FFC2FC4A6DA4F6E4FF4E" box="[928,1001,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="124" unit="mm" value="20.0">20 µm</quantity>
before porophore, and to about 
<quantity id="4C839B44A575FFC2FAB96DA4F091FF4E" box="[1363,1436,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="124" unit="mm" value="10.0">10 µm</quantity>
near the male pore, which opens on a narrowly cylindrical porophore (
<quantity id="4C839B44A575FFC2FC556C41F13BFEAA" box="[959,1078,260,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="5.0" metricValueMin="2.6" pageId="13" pageNumber="124" unit="mm" value="38.0" valueMax="50.0" valueMin="26.0">26–50 µm</quantity>
long, 
<quantity id="4C839B44A575FFC2FB6A6C40F1F8FEAA" box="[1152,1269,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="13" pageNumber="124" unit="mm" value="31.5" valueMax="33.0" valueMin="30.0">30–33 µm</quantity>
wide). Atrium length about 9–10 times porophore width, and 0.6–0.7 times body width. Two vasa deferentia per atrium, diameter 
<quantity id="4C839B44A575FFC2FF236C08F433FED2" box="[201,318,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="13" pageNumber="124" unit="mm" value="21.5" valueMax="24.0" valueMin="19.0">19–24 µm</quantity>
, length about 
<quantity id="4C839B44A575FFC2FE346C08F734FED2" box="[478,569,333,359]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="124" unit="mm" value="300.0">300 µm</quantity>
; posterior vas loops in XI, and both vasa join the atrial muscle layer near the middle, entering the lumen at about one third the distance from tip. (
<figureCitation id="13402A24A575FFC2FC7F6C34F10BFE3E" box="[917,1030,369,395]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="13" pageNumber="124">Fig. 7C,D</figureCitation>
)
</paragraph>
<paragraph id="8BC436A1A575FFC2FF2D6CD0F07EFDAE" blockId="13.[151,1437,152,791]" pageId="13" pageNumber="124">
Spermathecae in XI and XII, ampullae may extend into next segment (
<figureCitation id="13402A24A575FFC2FC1A6CD0F147FE1A" box="[1008,1098,405,431]" captionStart="FIGURE 7" captionStartId="11.[151,250,1725,1750]" captionTargetBox="[162,1421,391,1687]" captionTargetId="figure@11.[151,1436,381,1700]" captionTargetPageId="11" captionText="FIGURE 7. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior ends, external view of unmounted worms, both with pharynx partially everted. B. Chaetae; left to right: dorsal pair in IV, ventral chaeta in VI. C. Reproductive organs, segments X–XIII. D. Reproductive organs, segments IX–XI (second spermatheca was broken)." figureDoi="http://doi.org/10.5281/zenodo.3934003" httpUri="https://zenodo.org/record/3934003/files/figure.png" pageId="13" pageNumber="124">Fig. 7C</figureCitation>
). Ampullae irregular sac-like, epithelial layer about 
<quantity id="4C839B44A575FFC2FE636CFCF4C9FE66" box="[393,452,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="124" unit="mm" value="8.0">8 µm</quantity>
thick (
<figureCitation id="13402A24A575FFC2FDFB6CFCF765FE66" box="[529,616,441,467]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="13" pageNumber="124">Fig. 8E</figureCitation>
). Ectal ducts of spermathecae 
<quantity id="4C839B44A575FFC2FC506CFCF121FE66" box="[954,1068,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="13" pageNumber="124" unit="mm" value="75.0" valueMax="90.0" valueMin="60.0">60–90 µm</quantity>
long and up to 
<quantity id="4C839B44A575FFC2FB336CFDF02FFE66" box="[1241,1314,440,467]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="124" unit="mm" value="50.0">50 µm</quantity>
wide, usually tapered to about 
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near the pore; muscle layer thin (ca. 
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), lining cells somewhat columnar, about 
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high; lumen very narrow (
<figureCitation id="13402A24A575FFC2FE036F44F733FDAE" box="[489,574,513,539]" captionStart="FIGURE 8" captionStartId="12.[151,250,1818,1843]" captionTargetBox="[151,1433,381,1792]" captionTargetId="figure@12.[151,1436,381,1793]" captionTargetPageId="12" captionText="FIGURE 8. Eremidrilus gilita n. sp., from Gilita Creek. A. Anterior end of a whole-mounted worm. B. Anterior dorsal chaeta, from a whole-mount. C. Male porophore with ectal end of atrial duct, from a whole-mount. D. Atrium and porophore, from a sagittal section. E. Spermathecae from a sagittal section. F. Spermathecal duct, sagittal section. G. Nephridium; inset shows the ectal vesicle near the pore, from a whole mount. H. Surface of clitellum, with spermathecal pore in segment XI, close to intersegment 11/12. Note: specimens are facing left." figureDoi="http://doi.org/10.5281/zenodo.3934005" httpUri="https://zenodo.org/record/3934005/files/figure.png" pageId="13" pageNumber="124">Fig. 8F</figureCitation>
). Spermathecal pores without glands or distinct epidermal modifications.
</paragraph>
</subSubSection>
<subSubSection id="C361652AA575FFC2FF2D6F60F14EFCA2" pageId="13" pageNumber="124" type="discussion">
<paragraph id="8BC436A1A575FFC2FF2D6F60F14EFCA2" blockId="13.[151,1437,152,791]" pageId="13" pageNumber="124">
<emphasis id="B90FEAB3A575FFC2FF2D6F60F436FD8A" bold="true" box="[199,315,549,575]" pageId="13" pageNumber="124">Remarks.</emphasis>
The position of the spermathecal pores in line with the ventral chaetae, with the second pair close to intersegment 12/13, is similar to that of other 
<taxonomicName id="4C7B4D22A575FFC2FD756F0CF629FDD6" authorityName="Fend &amp; Rodriguez" authorityYear="2013" box="[671,804,585,611]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="genus">
<emphasis id="B90FEAB3A575FFC2FD756F0CF629FDD6" box="[671,804,585,611]" italics="true" pageId="13" pageNumber="124">Eremidrilus</emphasis>
</taxonomicName>
species having two spermathecal segments. Male pores open on a short porophore, narrower than that of 
<taxonomicName id="4C7B4D22A575FFC2FD286F2BF659FD32" authority="Fend &amp; Rodriguez, 2020" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[706,852,621,647]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="humboldti" status="sp. nov.">
<emphasis id="B90FEAB3A575FFC2FD286F2BF659FD32" box="[706,852,621,647]" italics="true" pageId="13" pageNumber="124">E. humboldti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57C8A575FFC2FCB16F28F697FD32" box="[859,922,621,647]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B90FEAB3A575FFC2FCB16F28F69EFD32" bold="true" box="[859,915,621,647]" pageId="13" pageNumber="124">n. sp</emphasis>
.
</taxonomicNameLabel>
and 
<taxonomicName id="4C7B4D22A575FFC2FC3B6F2BF19CFD32" baseAuthorityName="Cook" baseAuthorityYear="1971" box="[977,1169,621,647]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="allegheniensis">
<emphasis id="B90FEAB3A575FFC2FC3B6F2BF19CFD32" box="[977,1169,621,647]" italics="true" pageId="13" pageNumber="124">E. allegheniensis</emphasis>
</taxonomicName>
(see below), and markedly different in shape from the concave, annular porophore of 
<taxonomicName id="4C7B4D22A575FFC2FCB06FD7F6DDFD1E" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[858,976,658,683]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="artzaini">
<emphasis id="B90FEAB3A575FFC2FCB06FD7F6DDFD1E" box="[858,976,658,683]" italics="true" pageId="13" pageNumber="124">E. artzaini</emphasis>
</taxonomicName>
(a porophore is absent in 
<taxonomicName id="4C7B4D22A575FFC2FB1B6FD7F091FD1E" authority="Fend &amp; Rodriguez, 2020" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[1265,1436,658,683]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="montanensis" status="sp. nov.">
<emphasis id="B90FEAB3A575FFC2FB1B6FD7F091FD1E" box="[1265,1436,658,683]" italics="true" pageId="13" pageNumber="124">E. montanensis</emphasis>
</taxonomicName>
<emphasis id="B90FEAB3A575FFC2FF7D6FF0F5DAFD7A" bold="true" box="[151,215,693,719]" pageId="13" pageNumber="124">
<taxonomicNameLabel id="A23C57C8A575FFC2FF7D6FF0F5DAFD7A" box="[151,215,693,719]" pageId="13" pageNumber="124" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). The small size of the atrium and the thin atrial musculature most closely resemble those of 
<taxonomicName id="4C7B4D22A575FFC2FAE96FF3F09BFD7A" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[1283,1430,693,719]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="humboldti">
<emphasis id="B90FEAB3A575FFC2FAE96FF3F09BFD7A" box="[1283,1430,693,719]" italics="true" pageId="13" pageNumber="124">E. humboldti</emphasis>
</taxonomicName>
, but in that species the atrium is shorter and the porophore is relatively large (see 
<tableCitation id="C6F9031AA575FFC2FBCC6F9CF171FD46" box="[1062,1148,729,755]" captionStart="TABLE 2" captionStartId="16.[151,239,154,178]" captionText="TABLE 2. Selected measurements of Eremidrilus species with two spermathecal segments. Abbreviations: H, height; L," pageId="13" pageNumber="124">Table 2</tableCitation>
). The nephridial vesicles, observed in middle segments, appear more distinctive than those of 
<taxonomicName id="4C7B4D22A575FFC2FC7A6FBBF131FCA2" authorityName="Fend &amp; Rodriguez" authorityYear="2020" box="[912,1084,766,791]" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="montanensis">
<emphasis id="B90FEAB3A575FFC2FC7A6FBBF131FCA2" box="[912,1084,766,791]" italics="true" pageId="13" pageNumber="124">E. montanensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>