On Venezuelan pholcid spiders (Araneae, Pholcidae)
Author
Huber, Bernhard A.
33607F65-19BF-4DC9-94FD-4BB88CED455F
Zoological Research Museum Alexander Koenig, Bonn, Germany.
b.huber@leibniz-zfmk.de
Author
Villarreal, Osvaldo
679C385E-B068-4351-9D2F-97753E534C26
Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. & Museu Nacional / UFRJ, Rio de Janeiro, Brazil.
osvaldovillarreal@gmail.com
text
European Journal of Taxonomy
2020
2020-10-01
718
1
317
journal article
10.5852/ejt.2020.718.1101
4069574
F9E9A91E-488C-4DB1-9361-E788E9AC5BC1
Canaima avila
Huber
sp. nov.
urn:lsid:zoobank.org:act:
EF382001-C5C1-47E8-9679-E75E4C6F82A5
Figs 52–53
,
81–84
,
93–95
, 1034
Diagnosis
Distinguished from known congeners by armature of male chelicerae (
Fig. 83
; distinctive pair of distal processes and pair of low frontal humps), by shape of procursus (
Fig. 82
; distinctive subdistal dorsal widening – arrow in
Fig. 82
), by strongly banded legs (
Figs 52–53
; femora with 3–4 dark rings), and by external and internal female genitalia (
Figs 84
,
93
): strong lateral sclerites, small median sclerite; pair of internal anterior pockets/folds; elongate transversal pore plates close together; note that female of
C. merida
Huber, 2000
is unknown.
Etymology
The species name refers to the
type
locality; noun in apposition.
Type
material
VENEZUELA
–
Miranda
•
♂
holotype
,
ZFMK
(Ar 21832),
El Ávila National Park
,
near La Julia, ‘site 2’
(
10.5037° N
,
66.8107° W
),
1060 m
a.s.l.
, dry forest,
22–23 Feb. 2020
(
B.A. Huber
,
O. Villarreal M.
)
.
Other material examined
VENEZUELA
–
Miranda
•
4 ♂♂
,
5 ♀♀
,
ZFMK
(Ar 21833), and
8 ♀♀
, 7 juvs in pure ethanol,
ZFMK
(Ven20-179), same collection data as for holotype
.
Description
Male
(
holotype
)
MEASUREMENTS. Total body length 1.25, carapace width 0.60. Distance PME–PME
50 µm
; diameter PME
60 µm
; distance PME–ALE
30 µm
; AME absent. Leg 1: 10.9 (2.8 +0.2+2.8 +4.3+ 0.8), tibia 2: 1.6, tibia 3: 1.1, tibia 4: 1.3; tibia 1 L/d: 56.
COLOR (in ethanol). Prosoma and legs pale ochre-yellow, carapace and clypeus medially darker, legs with indistinct dark rings on femora (three) and tibiae (two); abdomen pale gray, dorsally light brown with large dark marks.
BODY. Habitus as in
Fig. 52
. Ocular area moderately raised. Carapace with distinct thoracic groove. Clypeus unmodified but slightly more protruding than in female. Sternum wider than long (0.44/0.32), with pair of small but distinct anterior processes. Abdomen globular.
CHELICERAE. As in
Fig. 83
, with distinctive pair of distal processes overhanging fangs and pair of low, light frontal humps.
PALPS. As in
Figs 81–82
; coxa with distinct retrolateral-ventral apophysis, trochanter barely modified, femur proximally with light retrolateral process, distally with small ventral process; procursus with distinctive subdistal dorsal widening (arrow in
Fig. 82
); genital bulb with complex distal process.
LEGS. Without spines and curved hairs; with higher than usual density of vertical hairs on tibiae (apparently only dorsally); retrolateral trichobothrium of tibia 1 at 11%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~13 pseudosegments.
Male
(variation)
Tibia
1 in
five males (incl.
holotype
): 2.7–2.9 (mean 2.8).
Female
Figs 81–84.
Canaima avila
Huber
sp. nov.
; from Miranda, El Ávila National Park (type locality; ZFMK Ar 21833).
81–82
. Left male pedipalp, prolateral and retrolateral views (arrow: distinctive dorsal widening of procursus).
83
. Male chelicerae, frontal view.
84
. Cleared female genitalia, dorsal view (arrows: anterior pockets/folds). Scale lines: 0.2 mm.
In general similar to male (
Fig. 53
) but without humps on sternum, clypeus less protruding, few vertical hairs on leg tibiae, sternum anteriorly laterally light brown, rings on legs more distinct, femora with four rings. Tibia
1 in
five females: 1.9–2.2 (mean 2.02). Epigynum (
Fig. 93
) barely protruding, anterior plate with strong lateral sclerites, medially whitish with small light brown sclerite; large light brown posterior plate. Internal genitalia (
Figs 84
,
94–95
) with elongate transversal pore plates close together and pair of unsclerotized internal anterior pockets/folds (arrows in
Fig. 84
); large median transparent receptacle.
Distribution
Known from
type
locality only, in
Venezuela
,
Miranda
(Fig. 1034).
Natural history
Most specimens were collected in masses of dead leaves and other debris suspended among lianas and other vegetation ~
1–2 m
above the ground.