Unraveling a new lineage of Hydrobiidae genera (Caenogastropoda: Truncatelloidea) from the Ponto-Caspian region Author Diana Delicado Author ladimir Pešić Author Peter Glöer text European Journal of Taxonomy 2016 208 1 29 journal article 10.5852/ejt.2016.208 2c5cb62e-9bc7-4fba-a4cb-e5a2fda8c719 2118-9773 269306 urn:lsid:zoobank.org:pub:F807AC79-2615-463B-8272-82523B46210A Genus Persipyrgula gen. nov. urn:lsid:zoobank.org:act:6E9922EE-56CD-4BBC-BF4E-9224C75859C0 Type species Pseudamnicola saboori Glöer & Pešić, 2009 . Diagnosis Shell ovate-conic, 2.5 to 4.5 mm high; large and convex body whorl; rest of the whorls small and slightly convex; aperture complete, pyriform, slightly angled on the top and separated from body whorl. Operculum corneous, yellowish, thin, pliable, ellipsoidal, paucispiral with submarginal nucleus. From one to two pairs of basal cusps in radular central tooth. Ctenidium occupying nearly the entire length of pallial cavity and bearing well-developed gill filaments. Osphradium opposite approximate middle of ctenidium. Bursa copulatrix posterior positioned relative to albumen gland; pigmented renal oviduct; one small pyriform seminal receptacle without duct. Prostate gland bean-shaped, about twice as long as wide; exit of the pallial vas deferens from the posterior-most section of the prostate gland and seminal vesicle entering the prostate gland in its middle section; penis simple, gradually tapering, with a distal end tapered. Nervous system with black pigmentation typically elongated. Etymology Referring to Persia, the historic name of the region where the genus was found, and to Pyrgula , type genus of the subfamily Pyrgulinae . Remarks Though similar in shell features, this genus differs from Pseudamnicola mainly due to its small and pyriform seminal receptacle in the female genitalia (long and elongate in Pseudamnicola ), shorter prostate gland, tapered and simple penis (triangular and folded in the latter), and because of the occasional possession of two pairs of basal cusps in the central radular tooth. Differences between Persipyrgula gen. nov. and Intermaria gen. nov. are: shell dimensions (larger in the latter), aperture not fused to body whorl in Persipyrgula gen. nov. , one (occasionally two) vs two pairs of basal cusps in the central radular tooth, small pyriform vs elongate seminal receptacle, and absence vs presence of a small distal lobe on the inner edge of the penis, respectively.