The fifth family of the true crickets (Insecta: Orthoptera: Ensifera: Grylloidea), Oecanthidae defin. nov.: phylogenetic relationships and divergence times Author Campos, Lucas Denadai De Author Dias, Pedro Guilherme Barrios De Souza Author Audino, Jorge Alves Author Desutter-Grandcolas, Laure Author Nihei, Silvio Shigueo text Zoological Journal of the Linnean Society 2023 2022-11-02 197 4 1034 1077 http://dx.doi.org/10.1093/zoolinnean/zlac066 journal article 10.1093/zoolinnean/zlac066 8c2036d0-8307-4b09-8c8b-c5eeea9d030a 0024-4082 7803427 4BB4333D-64F0-4485-9C2B-47546ECFE65F SUPERTRIBE PODOSCIRTIDI SAUSSURE, 1878 SUPERTRIB. NOV. Podoscirtites Saussure, 1878: 693 . Saussure, 1893: 257. Type genus: Podoscirtus Serville, 1838 . Distribution: Afrotropics, Australasia, Indo-Malaya, Oceania and Palaearctic. Figure 9. A, Ultratrella gracilis , male genitalia, ventral view. B, Taroba elephantina , apex of ovipositor: a, dorsal view, b, ventral view. C, Idiotrella karnyi , apex of ovipositor: a, dorsal view, b, ventral view. Scales: 1mm. Abbreviations: see Material and methods. Diagnosis: FWs and HWs well developed, frequently covering the entire abdomen, generally with stridulatory apparatus developed ( Fig. 10A ), sometimes with FWs bearing only longitudinal veins ( Fig. 10B ). When stridulatory apparatus developed, stridulatory vein (PCu) curved, forming a 90° angle, proximal branch parallel to CuPa ( Fig. 10C ); CuPa higher than CuPb in lateral view. TI inner tympanum, when present, generally profound. Male genitalia: MedLophi of pseudepiphallic sclerite when present, short; MedLophi dorsal to LLophi ( Fig. 10D ; Supporting Information, Fig. S8E ). Included tribes: Aphonoidini Gorochov, 1986, Podoscirtini Saussure, 1878 , Truljaliini Gorochov, 2020 defin. nov. Remarks: The clade corresponding in our phylogeny to the Podoscirtidi is clearly undersampled. Future phylogenetic studies on this supertribe, including more taxa, will be necessary to organize and verify more tribes as proposed herein. Besides, ProzƲenella bangaloriensis appears isolated in the ML (Supporting Information, Fig. S13 ) and MP (Supporting Information, Fig. S14 ) trees. In BI this taxon resolves inside Podoscirtini ( Fig. 3 ). Thus, we decided to maintain this genus as classified in Cigliano et al. (2022) . For these reasons, we avoid designing subtribes, as the relations between, and definitions of, present-day genus groups are still unclear.