<document ID-DOI="10.11646/zootaxa.5263.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804347" ID-ZooBank="FDB96626-95AF-4D4F-A6DD-FBC881C04D25" IM.taxonomicNames_approvedBy="valdenar" approvalRequired="2" approvalRequired_for_taxonomicNames="2" checkinTime="1680760719565" checkinUser="plazi" docAuthor="López, Guilherme E. L., Carrenho, Renan &amp; Schwertner, Cristiano F." docDate="2023" docId="03E887F8D0632732FF15FCEF8EB9FAEA" docLanguage="en" docName="zootaxa.5263.3.5.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyphuchus sculpturatus Stal 1860" docType="treatment" docVersion="3" lastPageNumber="406" masterDocId="FFD1FF80D0662737FF82FFAD8B3EFFCD" masterDocTitle="Over 170 years later: redescription of the genus Glyphuchus Stål (Hemiptera: Pentatomidae: Discocephalinae), with the first description of the male" masterLastPageNumber="410" masterPageNumber="401" pageNumber="406" updateTime="1681148365456" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title>Over 170 years later: redescription of the genus Glyphuchus Stål (Hemiptera: Pentatomidae: Discocephalinae), with the first description of the male</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>López, Guilherme E. L.</mods:namePart>
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<mods:affiliation>University of São Paulo, Museum of Zoology (MZUSP), Av. Nazaré, 481, CEP 04263 - 000, São Paulo, SP, Brazil. &amp; guilherme. enrique. lopez @ usp. br; https: // orcid. org / 0000 - 0002 - 6031 - 4453</mods:affiliation>
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<mods:namePart>Carrenho, Renan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4268-7309</mods:nameIdentifier>
<mods:affiliation>University of São Paulo, Museum of Zoology (MZUSP), Av. Nazaré, 481, CEP 04263 - 000, São Paulo, SP, Brazil. &amp; renan. carrenho @ gmail. com; https: // orcid. org / 0000 - 0003 - 4268 - 7309</mods:affiliation>
<mods:nameIdentifier type="email">renan.carrenho@gmail.com</mods:nameIdentifier>
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<mods:namePart>Schwertner, Cristiano F.</mods:namePart>
<mods:affiliation>University of São Paulo, Museum of Zoology (MZUSP), Av. Nazaré, 481, CEP 04263 - 000, São Paulo, SP, Brazil. &amp; Federal University of São Paulo, Department of Ecology and Evolutionary Biology, R. Prof. Artur Riedel, 275, CEP 09972 - 270, Diadema, SP, Brazil.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
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<mods:number>2023-04-06</mods:number>
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<mods:number>5263</mods:number>
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<mods:number>3</mods:number>
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<mods:end>410</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.3.5</mods:url>
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<heading bold="true" box="[151,585,833,861]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
<taxonomicName authority="Stal, 1860" authorityName="Stal" authorityYear="1860" box="[151,585,833,861]" class="Insecta" family="Pentatomidae" genus="Glyphuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis bold="true" box="[151,585,833,861]" pageId="5" pageNumber="406">
<emphasis bold="true" box="[151,455,834,861]" italics="true" pageId="5" pageNumber="406">Glyphuchus sculpturatus</emphasis>
<bibRefCitation author="Stal, C." box="[463,585,833,860]" pageId="5" pageNumber="406" pagination="1 - 84" refId="ref5765" refString="Stal, C. (1860) Bidrag till Rio Janeiro-Traktens Hemipter-Fauna. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 2, 1 - 84." type="journal article" year="1860">Stål, 1860</bibRefCitation>
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<taxonomicName authority="Stal, 1860: 17" authorityName="Stal" authorityPageNumber="17" authorityYear="1860" box="[151,555,903,927]" class="Insecta" family="Pentatomidae" genus="Glyphuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis box="[151,404,903,927]" italics="true" pageId="5" pageNumber="406">Glyphuchus sculpturatus</emphasis>
<bibRefCitation author="Stal, C." box="[411,555,903,927]" pageId="5" pageNumber="406" pagination="1 - 84" refId="ref5765" refString="Stal, C. (1860) Bidrag till Rio Janeiro-Traktens Hemipter-Fauna. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 2, 1 - 84." type="journal article" year="1860">Stål, 1860: 17</bibRefCitation>
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; 
<treatmentCitation author="Stal, C." box="[566,711,903,927]" page="12" pageId="5" pageNumber="406" year="1872">
<bibRefCitation author="Stal, C." box="[566,711,903,927]" pageId="5" pageNumber="406" pagination="1 - 159" refId="ref5890" refString="Stal, C. (1872) Enumeratio Hemipterorum. Bidrag till en f ˆ rteckning ˆ fver alla hittels kanda Hemiptera, jemte systematiska meddelanden. 2. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 10 (4), 1 - 159." type="journal article" year="1872">Stål, 1872: 12</bibRefCitation>
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; 
<treatmentCitation author="Kirkaldy, G." box="[722,929,903,927]" page="221" pageId="5" pageNumber="406" year="1909">
<bibRefCitation author="Kirkaldy, G." box="[722,929,903,927]" pageId="5" pageNumber="406" refId="ref4971" refString="Kirkaldy, G. (1909) Catalogue of the Hemiptera (Heteroptera). V. I: Cimicidae. Felix L. Dames, Berlin, xl + 392 pp." type="book" year="1909">Kirkaldy, 1909: 221</bibRefCitation>
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; 
<treatmentCitation author="Petrulevicius, J. F. &amp; Popov, Y. A." box="[940,1273,903,927]" page="24" pageId="5" pageNumber="406" year="2014">
<bibRefCitation author="Petrulevicius, J. F. &amp; Popov, Y. A." box="[940,1273,903,927]" pageId="5" pageNumber="406" pagination="23 - 33" refId="ref5299" refString="Petrulevicius, J. F. &amp; Popov, Y. A. (2014) First fossil record of Discocephalinae (Insecta. Pentatomidae): a new genus from the middle Eocene of Rio Pichileufu, Patagonia, Argentina. Zookeys, 422, 23 - 33. https: // doi. org / 10.3897 / zookeys. 422.6750" type="journal article" year="2014">Petrulevičius &amp; Popov, 2014: 24</bibRefCitation>
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, 28–30, 
<figureCitation box="[1360,1432,903,927]" captionStart="FIGURE 3" captionStartId="4.[152,255,1819,1843]" captionTargetBox="[151,1436,181,1765]" captionTargetId="figure-29@4.[151,1436,181,1795]" captionTargetPageId="4" captionText="FIGURE 3. Female holotype of G. sculpturatus deposited at NHRS. (A) dorsal view; (B) ventral view; (C) genital plates; (D) lateral view of the head, arrow indicating the curved labrum; (E) original labels. Scale bars: 1.0mm. Lettering: lt8, laterotergites 8; lt9, laterotergites 9; per, peritreme; vf8, valvifers 8. Photographed by Gunvi Lindberg (© 2022 Naturhistoriska riksmuseet). Original photos cropped, labelled, and reorganized." figureDoi="http://doi.org/10.5281/zenodo.7804355" httpUri="https://zenodo.org/record/7804355/files/figure.png" pageId="5" pageNumber="406">fig. 3E</figureCitation>
; 
<bibRefCitation author="Grazia, J. &amp; Campos, L. A. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F. &amp; Garbelotto, T. A. &amp; Roell, T. &amp; Barros, L. D." box="[199,390,934,958]" pageId="5" pageNumber="406" refId="ref4718" refString="Grazia, J., Campos, L. A., Fernandes, J. A. M., Schwertner, C. F., Garbelotto, T. A., Roell, T. &amp; Barros, L. D. (2023) Pentatomidae in Catalogo Taxonomico da Fauna do Brasil. PNUD. Available from: http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 2212 (accessed 14 February 2023)" type="url" year="2023">
Grazia 
<emphasis box="[273,324,934,958]" italics="true" pageId="5" pageNumber="406">et al.</emphasis>
, 2023
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.
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<subSubSection pageId="5" pageNumber="406" type="materials_examined">
<paragraph blockId="5.[151,1437,1004,1319]" pageId="5" pageNumber="406">
<materialsCitation collectionCode="NHRS" collectorName="F. Sahlb." country="Brazil" location="Brasil" pageId="5" pageNumber="406" specimenCount="1" typeStatus="holotype">
<typeStatus box="[151,255,1004,1030]" pageId="5" pageNumber="406">Holotype</typeStatus>
(by monotypy) female. Labels: 
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. // 
<collectorName box="[739,840,1004,1031]" pageId="5" pageNumber="406">F. Sahlb.</collectorName>
// Type. // NHRS-GULI / 000003136 (
<collectionCode box="[1294,1372,1004,1030]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="5" pageNumber="406">NHRS</collectionCode>
) (
<figureCitation captionStart="FIGURE 3" captionStartId="4.[152,255,1819,1843]" captionTargetBox="[151,1436,181,1765]" captionTargetId="figure-29@4.[151,1436,181,1795]" captionTargetPageId="4" captionText="FIGURE 3. Female holotype of G. sculpturatus deposited at NHRS. (A) dorsal view; (B) ventral view; (C) genital plates; (D) lateral view of the head, arrow indicating the curved labrum; (E) original labels. Scale bars: 1.0mm. Lettering: lt8, laterotergites 8; lt9, laterotergites 9; per, peritreme; vf8, valvifers 8. Photographed by Gunvi Lindberg (© 2022 Naturhistoriska riksmuseet). Original photos cropped, labelled, and reorganized." figureDoi="http://doi.org/10.5281/zenodo.7804355" httpUri="https://zenodo.org/record/7804355/files/figure.png" pageId="5" pageNumber="406">fig. 3E</figureCitation>
).
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<emphasis bold="true" box="[199,443,1076,1102]" pageId="5" pageNumber="406">Material Examined.</emphasis>
<materialsCitation collectingDate="2022-07-27" collectionCode="MZUSP" collectorName="E. B. Crispino" country="Brazil" county="Itatiaia" latitude="-22.452557" location="Casa do Pesquisador" longLatPrecision="1" longitude="-44.610306" municipality="Parque Nacional do Itatiaia" pageId="5" pageNumber="406" specimenCount="1" stateProvince="Rio de Janeiro">
<emphasis bold="true" box="[454,536,1076,1102]" pageId="5" pageNumber="406">
<collectingCountry box="[454,531,1076,1102]" name="Brazil" pageId="5" pageNumber="406">Brazil</collectingCountry>
:
</emphasis>
1♁ (
<collectionCode box="[601,700,1076,1102]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="406">MZUSP</collectionCode>
), 
<collectingRegion box="[723,896,1076,1102]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="406">Rio de Janeiro</collectingRegion>
, 
<collectingCounty box="[911,992,1076,1102]" pageId="5" pageNumber="406">Itatiaia</collectingCounty>
, 
<collectingMunicipality box="[1007,1328,1076,1103]" pageId="5" pageNumber="406">Parque Nacional do Itatiaia</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03E887F8D0632732FF15FCEF8EB9FAEA:8E9E6035D0632732FABDFB998A20FBBF" country="Brazil" county="Itatiaia" latitude="-22.452557" longLatPrecision="1" longitude="-44.610306" municipality="Parque Nacional do Itatiaia" name="Casa do Pesquisador" pageId="5" pageNumber="406" stateProvince="Rio de Janeiro">Casa do Pesquisador</location>
2, 
<geoCoordinate box="[322,475,1112,1138]" degrees="22" direction="south" minutes="27" orientation="latitude" pageId="5" pageNumber="406" precision="1" seconds="09.2" value="-22.452557">22°27’09.2”S</geoCoordinate>
<geoCoordinate box="[482,646,1112,1138]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="5" pageNumber="406" precision="1" seconds="37.1" value="-44.610306">44°36’37.1”W</geoCoordinate>
, 
<date box="[658,788,1112,1138]" pageId="5" pageNumber="406" value="2022-07-27">
<collectingDate box="[658,788,1112,1138]" pageId="5" pageNumber="406" value="2022-07-27">27.vii.2022</collectingDate>
</date>
, 
<collectorName box="[799,949,1112,1138]" pageId="5" pageNumber="406">E.B.Crispino</collectorName>
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.
</paragraph>
</subSubSection>
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<paragraph blockId="5.[151,1437,1004,1319]" box="[199,704,1148,1175]" pageId="5" pageNumber="406">
<emphasis bold="true" box="[199,373,1148,1174]" pageId="5" pageNumber="406">Redescription.</emphasis>
As per generic redescription.
</paragraph>
<paragraph blockId="5.[151,1437,1004,1319]" pageId="5" pageNumber="406">
<emphasis bold="true" box="[199,382,1184,1210]" pageId="5" pageNumber="406">Measurements.</emphasis>
Male (in mm, n = 1): total length 8.80; width of abdomen 5.70; head length 1.75; head width across eyes 2.25; interocular distance 1.35; interocellar distance 0.75; lengths of antennal segments: I 0.72, II 0.61, III 0.72, IV 0.84, V 0.97; labial segments lengths: I 0.75, II 1.25, III 0.95, IV 0.85; pronotum length along midline 1.90; pronotum width at humeral angles 4.75; scutellum length along midline 3.55; scutellum width at base 2.95.
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