Revision Of The Early Oligocene Flora Of Hrazený Hill (Formerly Pirskenberg) In Knížecí Near Šluknov, North Bohemia Author Kvaček, Zlatko Charles University in Prague, Faculty of Science, Albertov 6, CZ - 128 43 Praha 2, the Czech Republic; e-mail: kvacek @ natur. cuni. cz; Author Teodoridis, Vasilis Charles University in Prague, Faculty of Education, Magdalény Rettigové 4, CZ - 116 39 Praha 1, the Czech Republic; e-mail: vasilis. teodoridis @ pedf. cuni. cz; Author Zajícová, Jana Charles University in Prague, Faculty of Science, Albertov 6, CZ - 128 43 Praha 2, the Czech Republic; e-mail: jana. zajicova @ natur. cuni. cz. text Acta Musei Nationalis Pragae Series B 2015 2015-10-31 71 1 - 2 55 102 http://dx.doi.org/10.14446/amnp.2015.55 journal article 10.14446/AMNP.2015.55 2533-4069 13182943 Liriodendron haueri ETTINGSHAUSEN Pl. 3, Fig. 1–4, Pl. 6, Fig. 1 1869 Liriodendron haueri ETTINGSHAUSEN , p. 9, pl. 41, fig. 10–10b. 1961 Liriodendron procaccinii UNGER ; Knobloch, p. 273, pl. 7, fig. 1, 3. 1961 Styrax sp. ; Knobloch, p. 288, pl. 14, fig. 4. Leaves broadly ovate with four lobes arising from a point one third of the blade width or only shallowly bilobate, leaf blade 41–82 mm long and 21 – ca. 90 mm wide, base widely cuneate, petiolate, petiole maximum 50 mm long. Midrib strong and straight, secondary veins arising at an angle of 40–50°, opposite or alternate, every second joined by a broken tertiary vein to the next vein above and then looping. Lowermost pairs sending fine outer loops towards the margin. Tertiary venation forms polygonal fields. Mesophyll tissue with small lens-shaped secretory cells, cuticles smooth, the abaxial surface with scattered stomata openings, otherwise cell structure poorly preserved. D i s c u s s i o n. One aberrant leaf impression with only two shallow lobes was referred to Styrax by Knobloch (1961) but a similar leaf impression from Markvartice was assigned to Liriodendron on account of its epidermal anatomy ( Bůžek et al. 1976 , pl. 3, fig. 8). Of the available names for fossil species of Liriodendron (see e.g. Archenegg 1894 ) we prefer here Liriodendron haueri ETTINGSHAUSEN rather than L. procaccinii UNGER (selected by Knobloch 1958 , 1961 ) because the former name is based on a leaf impression from the Oligocene of North Bohemia (see Hably et al. 2001 , p. 27, pl. 20, fig. 2, Akhmetiev et al. 2009 , pl. 13, fig. 3) and was accepted by the previous authors dealing with other Palaeogene occurrences of Tulip tree foliage in this region (see e.g. Bůžek et al. 1976 , Walther 1998 ). L. procaccinii is a common designation for fossil foliage of Liriodendron distributed mainly in Europe during the late Neogene (see Saporta and Marion 1876 – Meximieux, Knobloch 1998 – Willershausen). L. haueri differs from L. procaccinii in acute lobes contrary to mostly rounded lobes in the Italian Neogene populations ( Knobloch 1998 , p. 14). Revision of the latter species type material from Senigallia, Italy ( Massalongo and Scarabelli 1859 , Kustatcher et al. 2014) is required. Two extant species differ in the surface sculpture of the abaxial cuticle. The leaves in L. tulipifera L. from E and SE North America are abaxially smooth, in L. chinense ( HEMSLEY ) SARGENT from eastern China and Vietnam are papillate. In this respect our material looks to be similar to L. tulipifera . It is noteworthy that in living species of Tulip tree the fruitlets survive in large quantities after the season under the trees while foliage readily decomposes over the winter. This is perhaps a reason why leaf impressions are less common than fruitlets in the fossil state (e.g., at Markvartice – Bůžek et al. 1976 , Roudníky – Kvaček et al. 2014 ). This is however not the case at the Knížecí site, where no fruitlets have yet been recovered. M a t e r i a l. Leaf impressions, NM-G2856a–f with cuticle, NM-G2859, NM-G2997, NM-G8591.