Rediscovery, redescription, and reclassification of the rare and unusual fly Pyrgometopa penicillata Kertész (Diptera: Drosophilidae) Author Pirani, Gabriela Laboratório de Evolução e Morfologia de Diptera, Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo, Av. Bandeirantes 3900, 14040 - 901, Ribeirão Preto, SP, Brazil. E-mail: gabi. pirani 3107 @ gmail. com; Author Grimaldi, David A. Curator, Division of Invertebrate Zoology; Professor, Gilder Graduate School, American Museum of Natural History, Central Park West at 79 th St, New York, NY. E-mail: grimaldi @ amnh. org. text Zootaxa 2019 2019-08-29 4661 3 445 456 journal article 25865 10.11646/zootaxa.4661.3.2 80f6b0f9-010f-4ae4-a944-fa8de3aa9a30 1175-5326 3446316 A166C53B-EC0F-427B-B6F4-1F39F92A3BD8 Genus Stegana Meigen Meigen, 1830: 79; Grimaldi (1990) revised diagnosis; Bachli et al. (2004) European species; Brake & Bachli (2008) world catalogue. Type species: S. nigra Meigen, 1830 ; Zetterstedt 1847: 2577 (= S. furta Linneaus, 1766 ). AF, AU, NE, NT, OR, PA. Pyrgometopa Kertész, 1901: 419 . Type species: P. penicillata Kertész . Monotypic. New Synonym . NT Diagnosis : Wing held against side of body at rest ( Fig. 2 , 7 ); apex bluntly pointed, veins R 4+5 and M 1 convergent, apices nearly touching, parallel at apex; C with row of minute, thorn-like spinules at apex ( Fig. 31–32 ). Mid tarsomeres 1–4 with 2 ventral rows minute, overlapping, scale-like setulae ( Figs. 34 ); hind tarsomeres 1–4 with one or two such rows ( Figs. 33, 34 ). Hypandrial complex very elaborate; phallapodeme usually long, fused to phallus and postgonites. Phallic guide usually present. Comments : Pyrgometopa has all the above-mentioned characters, fitting entirely in the definition of Stegana . The latter genus is cosmopolitan and more diverse in the tropical regions of the world. Furthermore, we have some undescribed species with modifications on the frons and ocellar tubercle that also fit entirely within Stegana and it is probable that they all form a clade within the genus.