Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae) Author Wanke, Dominic State Museum of Natural History Stuttgart, Entomology, Rosenstein & D- 70191 Stuttgart, Germany. & University of Hohenheim, Systematic Entomology (190 n), Garbenstr. 30, D- 70599 Stuttgart, Germany. Author Hausmann, Axel SNSB-ZSM, Bavarian State Collection of Zoology, Münchhausenstr. 21, D- 81247 Munich, Germany. Author Lee, Kyung Min Finnish Museum of Natural History, Zoology Unit, University of Helsinki, P. O. Box Author Murillo-Ramos, Leidys Grupo Biologi ́ a Evolutiva, Department of Biology, Universidad de Sucre, Sincelejo, Sucre, Colombia. Corresponding author. Author Sihvonen, Pasi Finnish Museum of Natural History, Zoology Unit, University of Helsinki, P. O. Box Author Rajaei, Hossein State Museum of Natural History Stuttgart, Entomology, Rosenstein & D- 70191 Stuttgart, Germany. text Zootaxa 2023 2023-10-24 5359 1 1 96 https://www.mapress.com/zt/article/download/zootaxa.5359.1.1/52094 journal article 10.11646/zootaxa.5359.1.1 1175-5334 10147920 11CC5175-7CCE-453F-88EB-3490E82F4972 Scopula marginepunctata terrigena 9 ♁/ ♀ , Iran , Zagros , Esfahan Umg. , Fereidun Shah , 2200 m , 9.vii.1999 , leg. A. Hofmann , J. Meineke ; 2 ♁/ ♀ , Iran , Zanjan , Takht –e Suleiman , Barbar Naza , 2462 m , 5.vii.2013 , N36°33’49.0’’ , E 47°17’57.7 ’, leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 1 ♁/ ♀ , Iran , Ardabil , Kuhha –ye Tales , Khal Khal W , Meragin vic., 2400 m , 1.viii.2006 , leg. A. Hofmann , J. Meineke ; 1 ♁/ ♀ , Iran , prov. Hamadan , Hamadan NNE, Razan N , Gardaneh Avaj , 2100–2150 m , 19.vi.1998 , leg. A. Hofmann , J.–U. Meineke ; 2 ♁/ ♀ , Iran , prov. Azarbayjan –e– Gharbi , Takht –e Suleyman , 20 km E, Gharawol–Khaneh , 2300–2500 m , 29./ 30.vi.2009 , leg. A. Hofmann , J.U.Meineke , H. Rajaei ; 1 ♁/ ♀ , Iran , prov. Azarbayjan –e– Shargi , Kuh –e Bozghush , Sarab S , 2233 m , 28.vi.2013 , N 37°45’49.4’’ , E 47°21’30.8’’ , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 2 ♁/ ♀ , Iran , prov. Azarbayjan –e– Shargi , Gharek Aghai , 2200 m , 4.vi.2013 , N 36°50’16.5’’ , E 46°58’07.5’’ , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 2 ♁/ ♀ , Iran , prov. Azarbayjan –e– Shargi , Sahand Mt. , S Batmanghelich S , 2358 m , 29.vi.2013 , N 37°34’29.8’’ , E 46°27’03.4’’ , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 2 ♁/ ♀ , Iran , Zanjan NE, Gard –ye Alamut N , 2000–2300 m , 1./2.vii.[20]09, leg. A. Hofmann , J.U. Meineke , A. Naderi , H. Rajaei ; 2 ♁/ ♀ , Iran , Zanjan , Zanjan E to Gill van, N 36°45’21.8’’ , E 48°49’20.7’’ , 1900 m , 6.vii.2013 , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 8 ♁/ ♀ , Iran , Golestan , Shahkuh 20 km W, Paband , 2000 m , 31.v.2008 , leg. J.–U. Meineke , W. Kramer ; 1 ♁/ ♀ , Iran , Golestan , Tange Gol , N 37°22’13.2’’ , E 55°56’10.2’’ , 700 m , 21.vi.2013 , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 7 ♁/ ♀ , Iran , Khorasan , Koppe Dag , 1200 m , Robat 20 km SW Izman Olia , 25./ 26.v.2008 , leg. J.–U. Meineke , W. Kramer ; 6 ♁/ ♀ , Iran , Khorasan –e Shomali , KopetDagh , Izman –e Sufla vic., N 37°51’31.9’’ , E 57°32’27.2’’ , 1823 m , 2.vi.2013 , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 2 ♁/ ♀ , Iran , Khorasan –e Shomali , Allah –o Akbar Mt. , 1763 m , 18.vi.2013 , N37°19’27.2’’ , E 58°43’26.1 ’, leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 2 ♁/ ♀ , Iran , Khorasan –e Shomali , Kopet–Dagh , Darreh Hersh , 2038 m , 19.vi.2013 , N37°40’23.2’’ , E 58°09’22.6 ’, leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 1 ♁/ ♀ , Iran , prov. Khorasan , Kuh –e Sorb , Kadkan S , 2200–2400 m , leg. A. Hofmann , J.U. Meineke ; 2 ♁/ ♀ , Iran , Esfahan , Semirom W, Pashmeh Kuh , Hochtal , 2800 m , 31.v.–2.vi.2004 , leg. A. Hofmann , J.–U. Meineke , W.G. Tremewan ; 1 ♁/ ♀ , Iran , Khorasan –e Razavi , Nishabour to Quochan , Bar vic., 1718 m , 17.vi.2013 , N36°30’27.9’’ , E 58°44’30.8 ’, leg. J.U. Meineke , H. Rajaei , B. Hafezi ; all in PCJM. 9 ♁/ ♀ , N– Iran , Amarlu östl., Rudbar , 1000 m , 27.ix.1970 , leg. G. Ebert , g. preps ( ♀ ) 0875, 0876/2020 D. Wanke; 1 ♁/ ♀ , N– Iran , Teheran / Evin , ca. 1400 m , 4. – 10.x.1970 , leg. G. Ebert ; 1 ♁/ ♀ , N– Iran , Talysch , Asalem , 1300 m , 29.ix.1970 , leg. G. Ebert ; 10 ♁/ ♀ , N– Iran , Elburs – Mts , Masandaran , Polur , Damavand , 11.vii.1972 , 2200 m , leg. Ebert & Falkner ; 8 ♁/ ♀ , N– Iran , Elburs Mts. , Masandaran , Polur , Damavand , 7.–10.vii.1972 , 2500 m , leg. Ebert & Falkner , (♁) g. prep. 1133/2021 D. Wanke; 56 ♁/ ♀ , N– Iran , Masandaran , Golestan – Wald , 60 km E Gonbad Qabus , 8.vii. 1972 , 510 m, leg. Ebert & Falkner , g. preps (♁) 0903/2020, 1132/2021 ( ♀ ) 0904/2020 D. Wanke; 26 ♁/ ♀ , NIran, E Alborz , Prov. Mazandaran , E Gorgan , S Aliabad , oberh. Shirinabad , N 36°47’21’’ , E 55°01’25’’ , 21.v.2005 , 1100 mNN, leg. R. Trusch , G. Petschenka & B. Müller , SMNK E. Lep. 215; 3 ♁/ ♀ , N– Iran , 10 km s. Chalus , 130 m , 26.v.1969 , leg. G. Ebert ; 3 ♁/ ♀ , N– Iran , Masandaran , Lar–Tal , NW Polur , 2200 – 2500 m , 20.vii.1975 , leg. Müller ; 1 ♁/ ♀ , Iran , prov. Mazandaran , Elburs Mts. , S Shah Kuh –e Bala , 2400 m , N 36°33’ , E 54°36’ , 20.vii.2003 , leg. G. Ebert & R Trusch (lux) ; 8 ♁/ ♀ , Persia sept., Elburs mts.c.s., Tacht i Suleiman , Särdab –Tal ( Vanderban ), 25–2700m , 14.–18.7. [19]37, leg. E. Pfeiffer & W. Forster , München , g. preps (♁) 0787, ( ♀ ) 0786/2020 D. Wanke; 2 ♁/ ♀ , Persia [ Iran ], Särdabtal , 1500 m , 19.–22.vii.[19]37; 2 ♁/ ♀ , Iran , prov. Esfahan , C– Zagros , 2600 m , 2 km NE of Semirom , 03.–04.vi.2005 , leg. P. Gyulai & A. Garai , g. prep. (♁) 0781/2020 D. Wanke; 3 ♁/ ♀ , Iran , prov. Hamadan , 5 km SW of Aradj Pass to Razan , 2500 m , 1. – 02.vi.2005 , leg. P. Gyulai & A. Garai ; 2 ♁/ ♀ , Iran , prov. Tehran , Elburz mts. 3 km NNW Shemshak , N36°02’ , E051°28’ , 24.vii.2003 , 2860 mNN, lux, leg. G. Ebert & R. Trusch ; 1 ♁/ ♀ , Nordiran , Elbursgebirge östl. Shemshak , 50 km nördl. Teheran , 2100 – 2500 m , 8.–24.vi.1973 , leg. G. Junge ; 3 ♁/ ♀ , NW– Iran , 17 km nw. Maku , 1400 m , 4.vi.1975 , leg. H. G. Amsel , g. prep. ( ♀ ) 1049/2021 D. Wanke; 1 ♁/ ♀ , N– Iran , Elburs Geb. , Polour , 1600 m , 21.vi.1969 , leg. H. G. Amsel ; 1 ♁/ ♀ N– Iran , Elburs – Mts. , Prov. Tehran , Arangeh 25 km N Karadj , 1550 m , 1.–6.vi.1972 , leg. Ebert & Falkner ; 1 ♁/ ♀ , N– Iran , Elburs Mts. , Prov. Tehran , 15 km E Gatschsar , 17.viii.1972 , 2800 m , leg. Ebert ; 2 ♁/ ♀ , N– Iran , 12 km n. Amarlu , 250 m , 28.x.1970 , leg. G. Ebert ; 1 ♀ , Iran , Tehran , Qollhak , 1400 m , 19.vi.1961 , leg. J. Klapperich , g. prep. 0784/2020 D. Wanke; 1 ♀ , same data, 04.vii.1961 , g. prep. 0785/2020 D. Wanke ; 1 ♁/ ♀ , Iran , 15.viii.1961 , leg. J. Klapperich ; 1 ♁/ ♀ , Iran , 10.vii.1961 , leg. J. Klapperich ; 1 ♁/ ♀ , Iran , 14.vii.1961 , leg. J. Klapperich ; 3 ♁/ ♀ , Iran , prov. Boyerahmad –va– Kohgiluyeh, SE – Zagros , 3000 m , 05.–06.vi.2005 , Kuh –e– Dena , n. Bijan pass, 6 km N of Cisakht , leg. P. Gyulai & A. Garai ; 2 ♁/ ♀ , Iran , prov. Azerbayejan, E – Sharqi , 10 km NW of Miyane , 14.–15.vi.2005 , leg. P. Gyulai & A. Garai ; 1 ♁/ ♀ , Iran , prov. Azerbayejan, E – Garbi , 8 km S of Shoet , 15.–16.vi.2005 , 1350 m , leg. P. Gyulai & A. Garai ; 49 ♁/ ♀ , Iran NE, Kopet Dagh , prov. Khorasan , ca. 50 km N Bojnurd , S Izmansufla , N 37°44’20’’ , E 57°25’53’’ , 1.240 mNN, 17.v.2005 , leg. Trusch , Petschenka , Müller , g. prep. (♁) 0840, 0906/2020 D. Wanke; 1 ♁/ ♀ , same data, 18.v.2005 ; 17 ♁/ ♀ , same data, 11.v.2005 ; 30 ♁/ ♀ , same data, 10.v.2005 ; 24 ♁/ ♀ , NE– Iran , Prov. Ostan –e Khorasan E Torbat de Heidarye S Fariman , Zharf vic. N 35°24’25.1’’ , E 59°56’08.2’’ 5.v.2008 , 2160 m , leg. R. Trusch , M. Falkenberg , B. Müller ; 31 ♁/ ♀ , same data, 6.v.2008 , g. prep. ( ♀ ) 0841/2020 D. Wanke ; 17 ♁/ ♀ , NE– Iran , Prov. Ostan –e Khorasan , Kopet Dagh , NW Mashad , N Tschenaran , N Radkan , Dolmeh Olia , N 36°55’56.6’’ , E 59°02’18.6’’ , 8.v.2008 , 1560 mNN, lux, leg. R. Trusch , M. Falkenberg & B. Müller , SMNK E– Lep. 247; 19 ♁/ ♀ , same data, 9.v.2008 ; 1 ♁/ ♀ , N– Iran , Elburs Mts. , N Teheran , Dizin Hotel , N 36°02’52.1’’ , E 51°24’58.1’’ , 2700 m , 20.vii.2006 , leg. R. Trusch ; 1 ♁/ ♀ , Iran – Centr. , Prov. Yazd , N Yazd , Chak Chak , N 32°20’ 07.8’’ , E 54°22’58.0’’ , 1.550 mNN, 10. – 11.iv.2007 , leg. R. Trusch , SMNK E– Lep. 234; 1 ♁, Iran , prov. Zanjan , Takhte Suleyman–Dandi road, 15 km E Takhte Suleyman , N36°36’23’’ , E47°21’02’’ , 2420 m , 29.vi.2009 leg. H. Rajaei , J.U. Meineke A. Hofmann , g. prep. 0828/2020 D. Wanke; 8 ♁/ ♀ , Iran , prov. Zanjan , 2350 m , W– Alborz range Tarom vic. 20 km NE of Zanjan , 13.–14.vi.2005 , leg. P. Gyulai & A. Garai E– Lep 221, g. prep. (♁) 0837/2020, 1146/2021 D. Wanke ; 3 ♁/ ♀ , NW– Iran , 12 km westl. Rezaiyeh , 1350 m , 3.v.1975 , leg. Amsel ; 1 ♁, N– Iran , Elburs Mts. , Prov. Tehran , 15 km E Gatschsar , 17.viii.1972 , 2800 m , leg. Ebert , g. prep. 1145/2021 D. Wanke; 1 ♁/ ♀ , same data, 5.viii.1972 , 2600 m ; 1 ♁, N– Iran , Elburs – Mts., S – Rand , Tehran – Evin , 24.vii.1972 , 1600 m , leg. G. Ebert , g. prep. 1139/2021 D. Wanke; 1 ♁/ ♀ , N– Iran , Shemshak , ca. 70 km n. Teheran , 20.vi.1969 , 1700 m , leg. H. G. Amsel ; all in SMNK. 6 ♁/ ♀ , Iran , Azerbaijan –e Gharbi prov. , Khoy to Ghotur road, Esteran vill., Alt. 1637 m , N 38°27’ 03.1’’ , E 44°44’33.6’’ , 1.vii.2013 , leg. H. Rajaei , J. U. Meineke , B. Hafezi , g. preps ( ♀ ) 0727, 0730/2020 D. Wanke; 2 ♁/ ♀ , Iran , Azerbaijan –e Gharbi prov. , Urmia to Seru road, after Eshke– Su , Alt. 1915 m , N 37°45’ 50.1’’ , E 44°48’46.4’’ , 30.vi.2013 , leg. H. Rajaei , J. U. Meineke , B. Hafezi ; 1 ♁/ ♀ , Iran , prov. Azarbayjan –e– Shargi , S Sahand Mt. , 2431 m , 30.vi.2013 , N 37°35’41’’ , E 46°27’18.4’’ , leg. J.U. Meineke , H. Rajaei , B. Hafezi ; 2 ♁/ ♀ , Iran , Zanjan prov. , E Zanjan , road to Gilvan , Alt. 1889 m , N36°45’21.8’’ , E48°49’20.7’’ , 6.vii.2013 , leg. H. Rajaei , J.–U. Meineke , B. Hafezi ; 2 ♁/ ♀ , Iran , Ghazwin–Alamut road, after Gardane Alamut , sandy road to Khanjar Bolagh village N 36°24’11’’ ; E50°12’52’’ , 2024 m , 1.–2.vii.2009 , H. Rajaei , J.–U. Meineke A. Hofmann ; 2 ♁/ ♀ , Iran , Khorasan Shomali , Babaaman , near Bojnurd , 26.vi.2016 , leg. Sh. Feizpour ; 1 ♁/ ♀ , Iran , Azerbaijan –e Gharbi prov. , 15 km W Jolfa , Sint– Stepanus Church , Alt. 951 m , N 38°58’ 43.2’’ , E 45°28’24.0’’ , 3.vii.2013 , leg. H. Rajaei , J. U. Meineke , B. Hafezi ; 4 ♁/ ♀ , Iran , Azerbaijan –e Sharghi prov. , S Ghareh Aghaj , after Argsnay–Sufla Alt. 2020 m , N 36°50’16.5’’ , E 46°58’07.5’’ , 4.vii.2013 , leg. H. Rajaei , J.U. Meineke , B. Hafezi ; 1 ♁, Iran , Lorestan , Dorud , Astaneh door, 1801 m , 33°24’48.1’’N, 49°08’42.5’’E, 25.vii.2016 , leg. Sh. Feizpour , g. prep. 0706/2020 D. Wanke; 1 ♁/ ♀ , Iran , Khorasan Razavi , 65 km Kalat road, Khor , 1431m , 36°38’15.1’’N, 59°54’04’’E, 16.vi.2016 , leg. Sh. Feizpour ; 1 ♁/ ♀ , Iran , Prov. Golestan , Shahrud – Golestan road, Shahkuh , 2585 m , 36°38’36’’N, 54°31’31’’E, 16.vii.2015 , leg. Sh. Feizpour ; 1 ♁, Iran , Mazandaran prov. , Noor , Sisangan Foresty Park , 35 m , N 36°32’09.3’’ , E 052° 04’55.8’’ , 24.vi.2013 , leg. H. Rajaei , J.U. Meineke , B. Hafezi , g. prep. 0725/2020 D. Wanke; 8 ♁/ ♀ , Iran , prov. Chaharmahal –va– Bakhtiyari , Dehnau , 2248m , 12.v.2010 , Leg. G. Petrányi, P . Hentschel ; 2 ♁/ ♀ , Iran , prov. Fars , Ardakan ( Sepidan ), 2400–2500m , 08–11.v.2010 , Leg. G. Petrányi , P. Hentschel ; 2 ♁/ ♀ , Iran , prov. Yazd , Sir Kuh , near Sanij , 2634m , 07.v.2010 , Leg. G. Petrányi , P. Hentschel ; 8 ♁/ ♀ , Iran , prov. Hamedan , Nehavand , 1855m , 13.v.2010 , Leg. G. Petrányi , P. Hentschel ; 12 ♁/ ♀ , Iran , prov. Golestan , Azad Sahr , 483m , 18.v.2010 , Leg. G. Petrányi , P. Hentschel ; 24 ♁/ ♀ , Iran , prov. Zanjan , Ab Bar , 1053m , 17.v.2010 , Leg. G. Petrányi , P. Hentschel ; 9 ♁/ ♀ , Iran , prov. Kordestan , Askaran , 1374m , 14–16.v.2010 , Leg. G. Petrányi , P. Hentschel ; 5 ♁/ ♀ , NW– Iran , Kaleibar , 1700m , 3.viii.1977 , leg. W. Thomas ; all in SMNS. 1 ♁/ ♀ , Nord – Iran , Schahkuh , West –abhg. Geröllzone , 1800–2000 m , Juni , Exp. Wernicke ; 1 ♁/ ♀ , Nordost – Iran , Kuh i Mirabi , Waldzone , 1600–1900m , Juli , Exp. Wernicke ; 2 ♁/ ♀ , Persia sept. [ Iran ], Elburs mts.c.s., Särdab Tal–Vandarban , 19–2200 m , 1.–3.vii.[19]37, E. Pfeiffer & W. Forster München leg.; all in ZFMK . 1 ♁/ ♀ , Iran , Lorestan prov. , Z. Zagros , W. Nar Abad , ca. 1900 m , 11.–20.v.2001 , leg. G. Müller ; 1 ♁/ ♀ , Iran , Golestan , Jahan Nama , 1800 m , 20.–21.x.2003 , 36’40.36,6/54’18.34,6, leg. Ch Wieser ; 1 ♁, Iran , Esfahan , 57 km S Borujen , 3 km S Dorahan , 31°36’N, 51°12’E, 8.vi.1997 , 1950 m , leg. A. Hofmann & P. Kautt , coll N. Pöll ; all in ZSM .