A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species Author Grichanov, Igor Ya. All-Russian Institute of Plant Protection, Podbelskogo 3, St Petersburg, Pushkin, 196608, Russia. grichanov@mail.ru text European Journal of Taxonomy 2023 2023-07-03 878 1 52 http://zoobank.org/07efe4b6-4184-41c7-82a5-747bf67ba093 journal article 57399 10.5852/ejt.2023.878.2153 0dfb60c4-1de3-4062-9443-15b55e7e0c88 2118-9773 8110550 07EFE4B6-4184-41C7-82A5-747BF67BA093 Thinophilus deemingi sp. nov. urn:lsid:zoobank.org:act: E987D48E-1FB1-4E86-801C-44BACFAB7907 Fig. 10 Diagnosis The small-sized Thinophilus deemingi sp. nov. is remarkable, having modified fore tarsus with basitarsus thick at apex; tarsal segments 2–4 short, triangular, with black lateral setae; segment 5 rounded, widened and flattened. As a whole, fore tibia ⅓ × as long as fore tarsus. In addition, wing crossvein dm-m ½ × as long as distal part of M 4. Other species of the Thinophilus indigenus group have practically simple fore tarsus, longer than fore tibia; wing crossvein dm-m as long as or ⅔ × as long as distal part of M 4 . Etymology The specific epithet is dedicated to the collector, the British dipterist Dr J.C. Deeming (NMWC) Material examined Holotype OMAN • ♂ ; “ Muscat , Qurum Beach ; [ 23°37′ N , 58°28′ E ]; 23 oct. 1990 ; J.С. Deeming leg.”; NMWC . Description Male ( Fig. 10A ) MEASUREMENTS . Body length 1.8 mm ; antenna length 0.5 mm ; wing length 1.8 mm ; wing width 0.6 mm . HEAD ( Fig. 10B ). Postcranium bluish black, whitish pollinose; frons, face and clypeus greenish black, grey pollinose; face under antennae about 2 × as wide as height of postpedicel; clypeus half as long as epistoma, 1.6 × as wide as long; palp yellow, bearing black bristly hairs; proboscis dark brown; 2 diverging ocellars; 1 vertical, 1 postvertical, stronger and longer than, and not in row with upper postoculars; upper postoculars uniseriate, black; middle and lower postoculars multiseriate, white, long; antennal scape, pedicel and postpedicel blackish dorsally, yellow ventrally ( Fig. 10C ); scape with scale-like inner projection; pedicel simple, convex on inner side; postpedicel apically browned, rounded, with short pubescence, as high as long; arista-like stylus dorsal, black and thick basally, thin distally, shortly pubescent; length ratio of scape to pedicel to postpedicel to stylus, 0.05/0.04/0.08/0.2. THORAX . Metallic, grey dusted; mesonotum greenish blue-black; pleura bluish black; no acrostichals; 6 dorsocentrals decreasing in length anteriorly; scutellum with 2 strong marginals and 2 minute laterals; few upper and lower, white propleural bristles of different length. LEGS . Fore coxa yellow; mid and hind coxae black, yellow at apex; legs mostly yellow; fore tarsomeres 2–5 and mid and hind tarsomeres 4–5 brown-black. FORE LEG . Coxa with black setae and bristles; femur ( Fig. 10D ) slightly thickened, with anteroventral row of about 6 black bristles, longer than femur height, with 3 preapical posteroventral setae; tibia bearing 2 anterodorsal, 2 posterodorsal and 3 apical short bristles; basitarsus thick at apex; tarsal segments 2–4 short, triangular, with black lateral setae ( Fig. 10E ); segment 5 rounded, widened and flattened; length of femur, tibia and tarsal segments (in mm): 0.68/0.55/0.18/0.05/0.05/0.04/0.09. MID LEG . Coxa with black setae; femur anteroventral row of black bristles, about as long as femur height, with 3 preapical posteroventral setae; tibia bearing 2 anterodorsal; 2 posterodorsal, 4 apical bristles; tarsal segment 5 weakly widened; length of femur, tibia and tarsal segments (in mm): 0.69/0.66/0.36/ 0.12/0.08/0.06/0.07. HIND LEG . Coxa with 1 black exterior bristle; femur with ventral row of several very short black setae, as long as anterodorsal setae; tibia bearing 2 anterodorsal, 2 posterodorsal bristles, 4 apicals; segment 5 weakly widened; length of femur, tibia and tarsal segments (in mm): 0.78/0.78/0.2/0.19/0.1/0.08/0.07. WING ( Fig. 10F ). Hyaline, without darker shades; veins brown-yellow, more yellowish at base; distal part of M 1+2 convex; tip of R 4+5 parallel with M 1+2 ; ratio of part of costa between R 2+3 and R 4+5 to that between R 4+5 and M 1+2 (in mm), 0.24/0.14; crossvein dm-m straight; ratio of dm-m to distal part of M 4 , 0.16/0.27; anal vein distinct; halter yellow; lower calypter yellow, with white cilia. ABDOMEN . Bluish black, weakly dusted; setae and hind-marginal bristles on tergites black, short; sternites bearing fungi ( Laboulbeniales ). Hypopygium ( Fig. 10G ) brown with yellow appendages; epandrial lobe fingerlike, with strong apical bristle; hypandrium short, apically concave; phallosoma massive, almost reaching apex of surstyli; phallus long and simple, strongly curved at apex of phallosoma, projected; surstylus straight (lateral view) with few longer bristles and several short apical and preapical setae ( Fig. 10H ); cerci dorsally fused almost to apex, boat-shaped, with marginal bristles ( Fig. 10G ). Female Unknown.