An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber Author Bartel, Christian Author Dunlop, Jason A. Author Giribet, Gonzalo text Zootaxa 2023 2023-05-29 5296 3 421 445 http://dx.doi.org/10.11646/zootaxa.5296.3.6 journal article 53499 10.11646/zootaxa.5296.3.6 2825cf24-234a-4b85-b997-673ee08019bf 1175-5326 7984131 59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A Species Leptopsalis breyeri sp. nov. Figs. 3–4 urn:lsid:zoobank.org:act: BE9130E7-26A1-470C-A505-C94D70DDDAD6 Holotype . MB .A.4457, ex coll. Jörg Wunderlich, F 2636. Type locality. Myanmar , Hukawng Valley; Burmese amber. Upper Cretaceous (Lower Cenomanian). Etymology. Named after Horst Breyer (the grandfather of Christian Bartel), who celebrated his 100 th birthday in January 2023 . This is a masculine patronym in the genitive. Diagnosis. Large and oval body, subtriangular in profile at the anterior end. Conical ozophores in type 2 position. Eyes possibly present. Chelicerae rather small and ornamented on median segment. Pedipalps thin and unmodified. Legs robust and granular. Tarsus IV with a long and fringed adenostyle near the middle. Opisthosomal sternal sulci mostly straight and parallel. Spiracles C-shaped. FIGURE 3. Leptopsalis breyeri sp. nov. , male holotype (MB.A.4457). A. Whole specimen, dorsolateral view. B. Same, ventral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ad, adenostyle; ap, anal plate; ch, chelicerae; fe, femur; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; tr, trochanter; legs numbered from I–IV. Scale bars equal 1 mm. FIGURE 4. Details of Leptopsalis breyeri sp. nov. , male holotype (MB.A.4457). A. Close-up of tarsus IV. B. Close-up of ventral side of opisthosoma. Abbreviations: ad, adenostyle; ap, anal plate; sp, spiracle; leg IV numbered. Scale bars equal 0.5 mm. Description. Body, total L 4.68, maximum prosomal W behind ozophores 1.64, maximum opisthosomal W 1.80 . Conical ozophores in type 2 position ( Fig. 3A, C ). L of ozophores 0.28, W at base 0.24, distance between base of ozophores 1.16. Possible eye lens present anterior to ozophores. Chelicerae relatively small and ornamented on median segment ( Fig. 3B, D ); basal segment L 0.51, median segment L 1.16; cheliceral fingers equivocal. Pedipalps thin, its tarsus sparsely covered with sensory setae; tarsus with a small claw on its tip; pedipalp length: fe 0.68, pa 0.32*, ti 0.40, ta 0.48. Legs robust, granular and with numerous setae on tarsus; tarsus IV bearing a relatively long and fringed adenostyle near the middle ( Fig. 4A ); leg tarsi not subdivided, bearing a single smooth claw; leg lengths: I tr 0.40, fe 1.28*, pa 0.44, ti 0.70, mt 0.32, ta 0.88*, total 4.02; II tr 0.48, fe 1.28, pa 0.44, ti 0.80, mt 0.36, ta 1.02, total 4.38; III tr 0.32, fe 1.00, pa 0.48, ti 0.80, mt 0.38, ta 0.72, total 3.70; IV tr 0.48, fe 1.20, pa 0.58, ti 0.82, mt 0.36, ta 1.12, total 4.56. Opisthosomal sternites completely granular. Sternites 2–3 fused to a subtriangular plate; sternites 4–8 free, their widths: 4, 2.06; 5, 1.87; 6, 1.69; 7, 1.44; 8, 1.13; 9, 0.81. Opisthosomal sternal sulci mostly straight and parallel. Gonostome equivocal. Spiracles C-shaped, maximum W 0.16 . Anal plate rounded and unmodified, W 0.53 , L 0.38 ( Fig. 4B ). Cuticle without conspicuous pits or depressions.