Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence
Author
Pyron, Robert Alexander
Author
Wallach, Van
text
Zootaxa
2014
3829
1
1
81
journal article
36820
10.11646/zootaxa.3829.1.1
a4b69269-5a4c-4b5b-9f89-6b314d3ba284
1175-5326
286556
75210CDC-AC6A-4624-A6F1-1BC969BC7CAA
Xerotyphlops
Hedges, Marion, Lipp, Marin & Vidal, 2014
Type
species.
Typhlops vermicularis
Merrem, 1820
Species content.
Xerotyphlops etheridgei,
Xer.
socotranus,
Xer.
vermicularis
,
and
Xer.
wilsoni
(?).
Diagnosis.
Xerotyphlops
can be distinguished from all other typhlopoids by the combination of the following characters: small- to moderate-sized (total length
92–405 mm
), stout- to moderate-bodied (length/width ratio 34–56) snakes with 20–30 scale rows (usually with reduction), 346–435 total middorsals, short to moderate tail (1.1–3.1% total length) with 7–13 subcaudals (length/width ratio 1.0–1.5), and apical spine. Dorsal and lateral head profiles bluntly rounded, narrow rostral (0.27–0.41 head width), inferior nasal suture in contact with second supralabial, preocular in contact with second and third supralabials, eye moderate with distinct pupil, T-III or T-V
SIP
, and postoculars 2–3. Lateral tongue papillae present; left lung absent, tracheal, cardiac and right lungs multicameral (with 22–29 + 5–9 + 2–9 chambers, respectively); testes segmented; hemipenis eversible, lacking retrocloacal sacs; and rectal caecum moderate (1.4–2.2% SVL).
Phylogenetic definition.
Includes the MRCA of
Xerotyphlops vermicularis
and
Xer. socotranus
and all descendants thereof, and all species more closely related to
Xer.
vermicularis
than to the
type
species of the 15 other typhlopid genera listed here.
Etymology.
Named after the desert (xeric) habitats of the species.
Distribution.
The
type
species (
Xerotyphlops vermicularis
) occurs from southeastern Europe through the Middle East. The other three species are more localized, with
Xer.
etheridgei
in
Mauritania
,
Xer. socotranus
on
Socotra
, and
Xer.
wilsoni
in southwestern
Iran
.
Remarks.
Assignment of
Xerotyphlops wilsoni
is tentative, as it is only known from the lost
holotype
, which has not been examined, though other putative specimens have been sighted near the
type
locality (B. Safaei, pers. comm.). Little morphological data is known for this species from the brief description (Table 2). This group likely contains additional cryptic species (
Kornilios
et al.
2012
).