<document id="3C77E1F6F06CA919093DF9A04A9688B8" ID-CLB-Dataset="235222" ID-DOI="10.11646/zootaxa.5251.1.1" ID-GBIF-Dataset="43227427-a867-4744-9e4c-2b2302524890" ID-ISSN="1175-5326" ID-Zenodo-Dep="7704190" ID-ZooBank="3326BF76-A2FB-4244-BA4C-D0AF81F55637" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678178331427" checkinUser="plazi" docAuthor="Guglielmone, Alberto A., Nava, Santiago &amp; Robbins, Richard G." docDate="2023" docId="03966A560F3DC73DBABF88B9B00EFB95" docLanguage="en" docName="zootaxa.5251.1.1.pdf" docOrigin="Zootaxa 5251 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5251.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Haemaphysalis humerosa Warburton &amp; Nuttall 1909" docType="treatment" docVersion="6" lastPageNumber="87" masterDocId="FFAF122E0F6BC76BBA288B32B56EFFCC" masterDocTitle="Geographic distribution of the hard ticks (Acari: Ixodida: Ixodidae) of the world by countries and territories" masterLastPageNumber="274" masterPageNumber="1" pageNumber="87" updateTime="1699476275219" updateUser="ExternalLinkService">
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<mods:title id="59117ADE245B18590FFD66DD3D29E456">Geographic distribution of the hard ticks (Acari: Ixodida: Ixodidae) of the world by countries and territories</mods:title>
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<mods:namePart id="C3B8DD8A1AAE09D426798EA3F43637BC">Guglielmone, Alberto A.</mods:namePart>
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<mods:namePart id="7C8827BC4C7B246D4AE6C9E51E88C3B8">Nava, Santiago</mods:namePart>
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<mods:namePart id="98155F78CEDE7A391E70B3CC9A6E9180">Robbins, Richard G.</mods:namePart>
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<emphasis id="B94B07520F3DC73DBABF88B9B5D5FC69" bold="true" box="[151,187,907,933]" italics="true" pageId="86" pageNumber="87">65.</emphasis>
<taxonomicName id="4C3FA0C30F3DC73DBA9788B8B677FC69" ID-CoL="3J7W8" ID-ENA="65643" authority="Warburton &amp; Nuttall, 1909" authorityName="Warburton &amp; Nuttall" authorityYear="1909" box="[191,793,906,933]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="humerosa">
<emphasis id="B94B07520F3DC73DBA9788B8B486FC69" bold="true" box="[191,488,906,933]" italics="true" pageId="86" pageNumber="87">Haemaphysalis humerosa</emphasis>
<bibRefCitation id="EFAEA6B10F3DC73DBBC688B9B677FC69" author="Warburton, C. &amp; Nuttall, G. H. F." box="[494,793,907,933]" pageId="86" pageNumber="87" pagination="57 - 76" refId="ref264310" refString="Warburton, C. &amp; Nuttall, G. H. F. (1909) On new species of Ixodidae, with a note on abnormalities observed in ticks. Parasitology, 2, 57 - 76. https: // doi. org / 10.1017 / S 0031182000001566" type="journal article" year="1909">Warburton &amp; Nuttall, 1909</bibRefCitation>
</taxonomicName>
.
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Australasian: 1) 
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, 2) 
<collectingCountry id="F3289BD00F3DC73DBE4B88B9B059FC69" box="[1123,1335,907,933]" name="Papua New Guinea" pageId="86" pageNumber="87">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="EFAEA6B10F3DC73DBF6B88B9B5BCFC05" author="Roberts, F. H. S." pageId="86" pageNumber="87" refId="ref247734" refString="Roberts, F. H. S. (1970) Australian ticks. CSIRO, Melbourne, 267 pp." type="book" year="1970">Roberts 1970</bibRefCitation>
, 
<bibRefCitation id="EFAEA6B10F3DC73DBAF5889DB40DFC05" author="Owen, I. L." box="[221,355,943,969]" pageId="86" pageNumber="87" pagination="1 - 163" refId="ref240497" refString="Owen, I. L. (2011) Parasites of animals in Papua New Guinea recorded at the National Veterinary Laboratory: a catalogue, historical review and zoogeographical affiliations. Zootaxa, 3143, 1 - 163. https: // doi. org / 10.11646 / zootaxa. 3143.1.1" type="journal article" year="2011">Owen 2011</bibRefCitation>
).
</paragraph>
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<paragraph id="8B80DB400F3DC73DBAEF88E1B713FBF9" blockId="86.[151,1437,906,1113]" pageId="86" pageNumber="87">
<bibRefCitation id="EFAEA6B10F3DC73DBAEF88E1B4AFFC21" author="Ernieenor, F. C. L. &amp; Ernna, G. &amp; Mariana, A." box="[199,449,979,1005]" pageId="86" pageNumber="87" pagination="387 - 400" refId="ref192121" refString="Ernieenor, F. C. L., Ernna, G. &amp; Mariana, A. (2017) Phenotypic and genotypic identification of hard ticks of the genus Haemaphysalis (Acari: Ixodidae) in Peninsular Malaya. Experimental and Applied Acarology, 71, 387 - 400. https: // doi. org / 10.1007 / s 10493 - 017 - 0120 - 3" type="journal article" year="2017">
Ernieenor 
<emphasis id="B94B07520F3DC73DBB1388E1B41CFC21" box="[315,370,979,1005]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
(2017)
</bibRefCitation>
allegedly found this tick in 
<collectingCountry id="F3289BD00F3DC73DB8D988E1B634FC21" box="[753,858,979,1005]" name="Malaysia" pageId="86" pageNumber="87">Malaysia</collectingCountry>
, but molecular differences with 
<emphasis id="B94B07520F3DC73DBE9388E1B04CFC21" box="[1211,1314,979,1005]" italics="true" pageId="86" pageNumber="87">bona fide</emphasis>
specimens of 
<taxonomicName id="4C3FA0C30F3DC73DBA9E88C5B4B9FBDD" authorityName="Warburton &amp; Nuttall" authorityYear="1909" box="[182,471,1015,1041]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="humerosa">
<emphasis id="B94B07520F3DC73DBA9E88C5B4B9FBDD" box="[182,471,1015,1041]" italics="true" pageId="86" pageNumber="87">Haemaphysalis humerosa</emphasis>
</taxonomicName>
indicate that the Malaysian specimens represent another species (
<bibRefCitation id="EFAEA6B10F3DC73DBE9688C5B0FEFBDD" author="Petney, T. N. &amp; Saijuntha, W. &amp; Boulanger, N. &amp; Chitimia-Dobler, L. &amp; Pfeffer, M. &amp; Eamudomkarn, C. &amp; Andrews, R. H. &amp; Ahamad, M. &amp; Putthasorn, N. &amp; Muders, S. V. &amp; Petney, D. A. &amp; Robbins, R. G." box="[1214,1424,1015,1041]" pageId="86" pageNumber="87" pagination="1 - 89" refId="ref243024" refString="Petney, T. N., Saijuntha, W., Boulanger, N., Chitimia-Dobler, L., Pfeffer, M., Eamudomkarn, C., Andrews, R. H., Ahamad, M., Putthasorn, N., Muders, S. V., Petney, D. A. &amp; Robbins, R. G. (2019) Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa, 4558, 1 - 89. https: // doi. org / 10.11646 / zootaxa. 4558.1.1" type="journal article" year="2019">
Petney 
<emphasis id="B94B07520F3DC73DBF3C88C5B023FBDD" box="[1300,1357,1015,1041]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
2019
</bibRefCitation>
), and this country is excluded from its range.
</paragraph>
<paragraph id="8B80DB400F3DC73DBAEF8F0DB00EFB95" blockId="86.[151,1437,906,1113]" box="[199,1376,1087,1113]" pageId="86" pageNumber="87">
<bibRefCitation id="EFAEA6B10F3DC73DBAEF8F0DB4FCFB95" author="Egan, S. L. &amp; Loh, S. M. &amp; Banks, P. B. &amp; Gillett, A. &amp; Ahlstrom, L. &amp; Ryan, U. M. &amp; Irwin, P. J. &amp; Oskam, C. L." box="[199,402,1087,1113]" pageId="86" pageNumber="87" pagination="1 - 13" refId="ref190534" refString="Egan, S. L., Loh, S. M., Banks, P. B., Gillett, A., Ahlstrom, L., Ryan, U. M., Irwin, P. J. &amp; Oskam, C. L. (2020) Bacterial community profiling highlights complex diversity and novel organisms in wildlife ticks. Ticks and Tick-borne Diseases, 11 (3) (article 101407), 1 - 13. https: // doi. org / 10.1016 / j. ttbdis. 2020.101407" type="journal article" year="2020">
Egan 
<emphasis id="B94B07520F3DC73DBB208F0DB42EFB95" box="[264,320,1087,1113]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
(2020)
</bibRefCitation>
found that more than one species may exist under the name 
<emphasis id="B94B07520F3DC73DBE1F8F0DB00EFB95" box="[1079,1376,1087,1113]" italics="true" pageId="86" pageNumber="87">
<taxonomicName id="4C3FA0C30F3DC73DBE1F8F0DB032FB95" authorityName="Warburton &amp; Nuttall" authorityYear="1909" box="[1079,1372,1087,1113]" class="Arachnida" family="Ixodidae" genus="Haemaphysalis" kingdom="Animalia" order="Ixodida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="humerosa">Haemaphysalis humerosa</taxonomicName>
.
</emphasis>
</paragraph>
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