The bats of the Congo and of Rwanda and Burundi revisited (Mammalia: Chiroptera) Author Cakenberghe, Victor Van 9A0E1AF5-C248-4648-9D64-443112890346 University of Antwerp, Department of Biology, Functional Morphology, Campus Drie Eiken, Universiteitsplein, 1, B- 2610 Antwerpen (Wilrijk), Belgium. & AfricanBats NPC, 357 Botha Ave, Kloofsig, 0157, Republic of South Africa. & urn: lsid: zoobank. org: author: 9 A 0 E 1 AF 5 - C 248 - 4648 - 9 D 64 - 443112890346 & Corresponding author: Victor. VanCakenberghe @ uantwerpen. be akenberghe@uantwerpen.be Author Tungaluna, Guy-Crispin Gembu 2AE2F96F-A138-419B-B650-6DB44D535D14&4C40421C-1D84-4BD1-A444-360300ECBEEC Faculté des Sciences, Université de Kisangani, Democratic Republic of the Congo. & Email: gembuguycrispin @ gmail. com & urn: lsid: zoobank. org: author: 2 AE 2 F 96 F-A 138 - 419 B-B 650 - 6 DB 44 D 535 D 14 & Faculté des Sciences, Université de Kisangani, Democratic Republic of the Congo. & Email: prescottmusaba @ yahoo. fr & urn: lsid: zoobank. org: author: 4 C 40421 C- 1 D 84 - 4 BD 1 - A 444 - 360300 ECBEEC gembuguycrispin@gmail.com&prescottmusaba@yahoo.fr Author Akawa, Prescott Musaba Author Seamark, Ernest B60CA9A1-D288-468E-AEEB-D0F136E96F5B AfricanBats NPC, 357 Botha Ave, Kloofsig, 0157, Republic of South Africa. & Centre for Wildlife Management, University of Pretoria, Private Bag X 20 Hatfield, Pretoria 0028, Republic of South Africa. & Email: ernest. seamark @ africanbats. org & urn: lsid: zoobank. org: author: B 60 CA 9 A 1 - D 288 - 468 E-AEEB-D 0 F 136 E 96 F 5 B ernest.seamark@africanbats.org Author Verheyen, Erik 86B40463-E3D9-4147-9ED3-D7302E0D64B6 Royal Belgian Institute of Natural Sciences - OD Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. & University of Antwerp, Department of Biology, Evolutionary Ecology, Campus Drie Eiken, Universiteitsplein 1, B- 2610 Antwerpen (Wilrijk), Belgium. & Email: erik. verheyen @ naturalsciences. be & urn: lsid: zoobank. org: author: 86 B 40463 - E 3 D 9 - 4147 - 9 ED 3 - D 7302 E 0 D 64 B 6 erik.verheyen@naturalsciences.be text European Journal of Taxonomy 2017 2017-12-18 382 1 327 journal article 21810 10.5852/ejt.2017.382 ea406606-dfbd-4121-9d8c-108041a032b4 2118-9773 3860077 FA508A12-9BDB-4A2B-9B0C-98FDD161443C Myonycteris torquata (Dobson, 1878) DEMOCRATIC REPUBLIC OF THE CONGO: 4 ?? [3 AL, 1 SK], unknown locality , unknown date, leg. Prescott Musaba, between 1 May 2010 and 31 May 2010 , leg. Nicolas Nesi, Blaise Kadjo and Christiane Denys (RMCA: a1.097-M-3194 to a1.097-M-3196, MNHN: ZM-2014-1168); 1? [UN], Albert National Park [= Virunga National Park] , unknown date, leg. unknown collector (INPBC: V1694.2); 1 ♂ [SS], Bakwakenge [= Bakwa-Kenge] , 3 Feb. 1980 , leg. Betu Nsankulu (RMCA: 80.020-M-0005); 4 ♀♀ , 2 ♂♂ [1 SS, 5 SK], Baliko , 11 Jun. 2012 , 12 Jun. 2012 , 13 Jun. 2012 , 14 Jun. 2012 , leg. William (“Bill”) T. Stanley (FMNH: 219592, 219593, 219680, 219594 to 219596); 2 ♂♂ [2 SS], Bena Mbala , 21 Feb. 1984 , leg. Betu Nsankulu (RMCA: 84.035-M-0004, 84.035-M- 0006); 6 ♂♂ , 4 ?? [10 AL], Boende , between 1 Jan. 1974 and 31 Dec. 1974 , between 1 Jan. 1967 and 31 Dec. 1967 , leg. P. Lootens (RMCA: 38650, 38651, 38671 to 38676, 75.035-M-0002, 75.035-M- 0003); 4 ♀♀ , 5 ♂♂ [9 UN], Bomane [= Bomaneh] , 21 May 2010 , 30 May 2010 , leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2127, CRT2130, CRT2144, CRT2156, CRT2157, CRT2195, CRT2196, CRT2200, CRT2078); 1 ♂ [SS], Cinputu , 13 Sep. 1983 , leg. Betu Nsankulu (RMCA: 84.035-M-0005); 2 ♂♂ [2 UN], Congo-Nil-Aka , 20 May 1952 , leg. H. de Saeger et al. , 20 May 1952 , leg. H. de Saeger (RBINS: 13525, RMCA: 13525); 14 ad ♀♀ , 17 ad ♂♂ , 15 sad ♀♀ , 14 sad ♂♂ , 38 ♀♀ , 40 ♂♂ [1 AL, 60 SK, 77 UN], Djabir , unknown date, 8 May 2011 , 9 May 2011 , 10 May 2011 , 12 May 2011 , leg. Prescott Musaba, 8 May 2011 , 9 May 2011 , 10 May 2011 , 11 May 2011 , 12 May 2011 , 13 May 2011 , leg. Prescott Musaba and Jonathan Kosele (RMCA: a1.097-M-3248, a1.097-M-3249, a1.097-M-3265 to a1.097-M-3268, a1.097-M-3282, a1.097-M-3250 to a1.097-M-3264, UNIKIS: DJB083, DJB084, DJB087, DJB088, DJB091, DJB092, DJB094, DJB097, DJB098, DJB102, DJB104, DJB106, DJB109, DJB111, DJB122, RMCA: a1.097-M-3269 to a1.097-M-3281, UNIKIS: DJB127, DJB128, DJB130, DJB133, DJB139, DJB142, DJB147, DJB154 to DJB167, DJB170, RMCA: a1.097-M-3223 to a1.097-M-3230, UNIKIS: DJB001, DJB003, DJB006, DJB008 to DJB011, DJB014, DJB016, DJB018 to DJB020, DJB022 to DJB024, DJB026, DJB027, DJB029, DJB030, DJB032, DJB034, DJB036, DJB037, RMCA: a1.097-M-3231 to a1.097-M-3247, UNIKIS: DJB040, DJB044, DJB046, DJB048 to DJB050, DJB052, DJB053, DJB055, DJB056, DJB058, DJB060, DJB062, DJB067, DJB070, DJB071, DJB169, DJB172); 5 ♀♀ , 2 ♂♂ [7 UN], Epulu , 29 Aug. 2015 , 30 Aug. 2015 , leg. André Malekani (UNIKIS: EPLU660, EPLU669, EPLU670, EPLU683, EPLU684, EPLU695, EPLU696); 1 imm ♂ , 1 ♀ [2 UN], Gangala-na-Bodio , 11 May 1948 , leg. unknown collector (RBINS: 11657, RMCA: 11657); 4 ♀♀ , 16 ♂♂ [20 UN], Île Mbiye [= Mbiye Island] , 16 Jan. 2012 , 25 Feb. 2012 , 30 Mar. 2012 , 31 Mar. 2012 , leg. Prescott Musaba, Tembele and Marc Boketshu, 19 Apr. 2014 , 21 Apr. 2014 , leg. Prescott Musaba, Moïse Bipoo, Richard Tamaru, Pauline Isude and Joseph Kussa, 7 Aug. 2015 , 28 Dec. 2015 , 8 Jan. 2016 , leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (UNIKIS: CHIM07, CHIM07/14, CHIM011/14, CHIM012/14, CHIM29, ABCHIM07,ABCHIM08, CHIM34, CHIM35, CHIM40 to CHIM45, CHIM47, CHIMA38, CHIMTB01, ABCHIM09, ABCHIM11); 1 ♀ [SA], Irangi , 11 Feb. 1984 , leg. Heinz Stephan (SMF: 65030); 2 ♀♀ [2 UN], Isai River , between 16 Dec. 1974 and 23 Dec. 1974 , leg. Michael D. Gallagher (HZM: 1.7798 , 2.7799 ); 1 ♀ [UN], Kamikoni , 17 May 1960 , leg. unknown collector (RBINS: 1694); 1 imm ♀ [UN], Karambi , 4Apr. 1992 , leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24567); 1 ♀ [SO], Kimpoko [= Kimpoko-Nsele] , 7 Nov. 1978 , leg. Tshinyenye (RMCA: 79.003-M-0001); 1 ♂ [UN], Kingabwa , 31 Dec. 1975 , leg. Tshinyenye (RMCA: 76.010-M-0004); 1 imm? [UN], Kinshasa , 2 May 1964 , leg. Jo Van Orshoven (ZMA: MAM.11163); 21 ?? [21 UN], Kinshasa City Province , 12 Feb. 2001 , between 1 Apr. 1999 and 30 Apr. 1999 , leg. unknown collector (SMF 92979 to 92983, 93281 to 93296); 1 ♀ , 1 ♂ [2 AL], Kisangani , between 1 Jun. 1982 and 30 Jun. 1982 , leg. Marc Colyn (RMCA: 82.031-M-0012, 82.031-M-0013); 2 ♂♂ [1 SK, 1 SO], Kitangua , 24 Dec. 1981 , leg. Betu Nsankulu (RMCA: 83.025-M- 0006, 83.025-M-0019); 20 ♀♀ , 8 ♂♂ , 1? [13 SK, 16 UN], Kona , unknown date, leg. Prescott Musaba, 12 May 2010 , 13 May 2010 , 14 May 2010 , 15 May 2010 , between 12 May 2010 and 13 May 2010 , leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (RMCA: a1.097-M-3160, UNIKIS: CRT1910, CRT1911, CRT1920, CRT1926, CRT1937, CRT1942 to CRT1945, CRT1947 to CRT1950, CRT1953, CRT1960, CRT1963, CRT1966 to CRT1972, CRT1979 to CRT1982, CRT1918); 3 ?? [3 AL], Konyola , prior to 5 Aug. 1988 , leg. Luc De Vos, Kimbembi and Alain Vandelanoote (RMCA: 88.025-M-0005 to 88.025-M-0007); 2 ♂♂ [2 UN], Kumbi , 7 Dec. 1993 , leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24896, MAM.24897); 13 ♀♀ , 14 ♂♂ , 1? [1 SK, 27 UN], Lieki , 21 May 2010 , 30 May 2010, 1 Jun. 2010 , 2 Jun. 2010 , 3 Jun. 2010 , leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2029, CRT2030, CRT2032 to CRT2035, CRT2037, CRT2282 to CRT2288, CRT2290 to CRT2296, CRT2057, CRT2021 to CRT2023, CRT2028, CRT2050, CRT2051); 1 ad ♂ [UN], Likandi River [= Litaki River] , 18 Apr. 1906 , leg. The Alexander-Gosling Expedition (BMNH: 1907.7.8.25 [ holotype Myonycteris wroughtoni K. Andersen, 1908 ]); 1 ♂ [UN], Likati River , 18 Apr. 1906 , leg. unknown collector (BMNH: 1907.7.8.26); 14 ?? [14 SS], Lovanium University [= Kinshasa] , 10 Jan. 1962 , 13 Jan. 1962 , 2 May 1962 , leg. De Bont, 22 Nov. 1962 , 3 Dec. 1962 , 20 Feb. 1963 , 17 Mar. 1963 , 23 Mar. 1963 , 30 Mar. 1963 , 2 Apr. 1963 , 19 Apr. 1963 , 26 Dec. 1966 , between 30 Mar. 1963 and 1 Apr. 1963 , between 1 Nov. 1962 and 1 Dec. 1962 , leg. De Bont and Jo Van Orshoven (RMCA: 31197, 31198, 32606, 32608, 32611, 31199, 32609, 32602, 32607, 32601, 32604, 32605, 32610, 32603); 1 ♂ [AL], Lubumbashi , 12 Oct. 2009 , leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-859); 1 ♂ [SS], Lukonga , 7 Oct. 1978 , leg. Betu Nsankulu (RMCA: 79.022-M-0006); 12 ?? [3 SS, 9 AL], Luluabourg [= Kananga] , 1 May 1964, 7 Nov. 1964 , 13 Feb. 1965 , 20 Feb. 1965 , 4 Mar. 1965 , 1 Apr. 1965 , 2 Apr. 1965 , prior to 1 Jun. 1965 , leg. Antoon Emeric Marcel De Roo (RMCA: 33336, 33338, 33413, 33414, 32660, 33333, 33339, 33340, 33334, 33335, 33332, 33337); 1 ♂ [UN], Lwana River , unknown date, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24568); 2 sad ♂♂ , 1 juv ♀ , 27 ♀♀ , 45 ♂♂ [4AL, 62 SK, 9 UN], Masako , unknown date, 27 Nov. 1987 , 28 Nov. 1987 , 29 Nov. 1987 , 30 Nov. 1987 , 16 Dec. 1987 , 17 Dec. 1987 , 22 Apr. 1988 , 25 Aug. 1989 , 29 Aug. 1989 , 30 Aug. 1989 , 24 Nov. 1989 , 22 Mar. 1990 , 25 Mar. 1990 , leg. Ndey B. Ifuta, 10 Jun. 2015 , 11 Jun. 2015 , 13 Jun. 2015 , leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (RMCA: 93.079-M-0278, UNIKIS: CHMS062, CHMS063, CHMS065, CHMS066, CHMS070 to CHMS072, CHMS075, CHMS077, RMCA: 93.079-M-0043 to 93.079-M-0051, 93.079-M-0287 to 93.079-M-0291, 93.079-M-0052 to 93.079-M-0059, 93.079-M-0228, 93.079-M-0292, 93.079-M-0276, 93.079-M-0272, 93.079-M-0275, 93.079-M-0277, 93.079-M-0270, 93.079-M-0026, 93.079-M-0027, 93.079-M-0214 to 93.079-M-0216, 93.079-M-0028 to 93.079-M-0033, 93.079-M-0217 to 93.079-M- 0219, 93.079-M-0267, 93.079-M-0273, 93.079-M-0034, 93.079-M-0220, 93.079-M-0284, 93.079-M- 0274, 93.079-M-0035 to 93.079-M-0042, 93.079-M-0221 to 93.079-M-0226, 93.079-M-0285, 93.079- M-0286); 2 ad ♀♀ , 1 ad ♂ , 1 imm ♀ [4 UN], Medje , 16 Apr. 1910 , 6 Sep. 1910 , 4 Apr. 1914 , leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48752, 48755, 48753, 48754); 3 ♂♂ [3 AL], Pandé, Forest N of , 11 Oct. 2009 , leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-856 to ZM- 2011-858); 6 ♀♀ [1 SS, 1 AL, 4 SK], Quatorze , 15 Jun. 2012 , 25 Jun. 2013 , 29 Jun. 2013 , 30 Jun. 2013 , leg. William (“Bill”) T. Stanley (FMNH: 219597, 222421, 222622, 222422 to 222424); 6 ♀♀ , 3 ♂♂ [9 UN], Rubi-Tele , 18 Aug. 2013 , 19 Aug. 2013 , 20 Aug. 2013 , 22 Aug. 2013 , 27 Aug. 2013 , leg. Guy- Crispin Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: RBTL502, RBTL503, RBTL516, RBTL520, RBTL529, RBTL530, RBTL541, RBTL542, RBTL575); 2 ♀♀ , 1? [1 SK, 2 UN], Sciere Forest , between 24 Nov. 1974 and 25 Nov. 1974 , leg. Michael D. Gallagher (HZM: 3.7866 to 5.7868); 1 ♀ , 2 ♂♂ [3 SS], Topetope , 10 Jun. 1966 , 5 Jul. 1966 , 10 Jul. 1966 , leg. Alexandre Prigogine (RMCA: 34381, 34383, 34382); 3 ♂♂ [3 UN], Uma , 10 Apr. 2014 , 12 Apr. 2014 , 13 Apr. 2014 , leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: UMA501, UMA624, UMA635); 14 ?? [14 AL], Wafanya [= Wafania] , 3 Dec. 1988 , 2 Sep. 1991 , 14 Sep. 1991 , 3 Oct. 1991 , 18 Apr. 1992 , 12 May 1992 , 29 May 1992 , between 1 Nov. 1991 and 30 Nov. 1991 , between 1 Apr. 1991 and 30 Apr. 1991 , between 1 Jan. 1991 and 31 Dec. 1991 , leg. P. Lootens (RMCA: 92.079-M-0068, 91.076-M-0082, 92.079-M-0065, 92.079-M-0070, 91.076-M-0065, 92.079-M-0066, 92.079-M-0069, 91.076-M-0098, 91.076-M-0100, 89.020-M-0039, 92.079-M-0063, 91.076-M-0020, 92.079-M-0064, 92.079-M-0067); 41 ♀♀ , 73 ♂♂ [37 SK, 77 UN], Yaekela , 2 May 2010 , 3 May 2010 , 4 May 2010 , 5 May 2010 , 6 May 2010 , 7 May 2010 , between 5 May 2010 and 13 May 2010 , between 5 May 2010 and 12 May 2010 , leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2305, CRT2307, CRT2316 to CRT2318, CRT2320, CRT2351 to CRT2360, CRT2362, CRT2365, CRT2366, CRT2369 to CRT2373, CRT2375 to CRT2383, CRT2385, CRT1801, CRT1802, CRT1804 to CRT1808, CRT1810, CRT1811, CRT1815, CRT1819, CRT1820, CRT2391, CRT2393, CRT2394, CRT2397 to CRT2400, CRT1824 to CRT1830, CRT1833 to CRT1840, CRT1842 to CRT1845, CRT1847, CRT1848, CRT1850 to CRT1853, CRT1855, CRT1857, CRT1859 to CRT1866, CRT1869 to CRT1878, CRT1883 to CRT1887, CRT1889 to CRT1896, CRT1831, CRT1832, CRT1841); 1 ♀ , 2 ♂♂ [2 SS, 1 AL], Yalosemba , 18 Jun. 1979 , 19 Jun. 1979 , leg. C. Brian Robbins, 23 Jun. 1979 , leg. Frieder Meier (USNM: 537694, 537695, MHNG: 1871.014); 3 ♀♀ , 11 ♂♂ , 1? [3 AL, 1 SK, 11 UN], Yoko , unknown date, 4 Sep. 2012 , 7 Sep. 2012 , leg. Prescott Musaba, 18 May 2009, 23 Sep. 2009 , 24 Sep. 2009 , 28 Sep. 2009 , leg. unknown collector, 4 Sep. 2012 , 5 Sep. 2012 , 6 Sep. 2012 , leg. Prescott Musaba, Tembele and Marc Boketshu (RMCA: a1.097-M-3161, UNIKIS: GE246, GE281, GE284, GE293, RMCA: a1.097-M-3192, a1.097-M-3193, UNIKIS: YK005CH, YK010CH, YK013CH, YK015CH, YK018CH, YK019CH, YK025CH, RMCA: a1.097-M-3191); 2 ♂♂ [2 UN], Zookis [= Kisangani] , 24 Dec. 2015 , leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (UNIKIS: ZCH05, ZCH12).