Two sea anemones (Cnidaria: Anthozoa: Actiniaria) from the Southern Ocean with evidence of a deep-sea, polar lineage of burrowing sea anemones
Author
Gusmão, Luciana C.
Author
Rodríguez, Estefanía
text
Zoological Journal of the Linnean Society
2021
2021-02-18
193
1392
1415
https://doi.org/10.1093/zoolinnean/zlaa176
journal article
3130
10.1093/zoolinnean/zlaa176
f0ab84e8-4ff4-46a8-a567-a9acd016305c
0024-4082
5752621
D000BAA9-51C4-46FE-AC9B-BB361B601FA2
FAMILY
HALCAMPOIDIDAE
APPELLÖF, 1896
Diagnosis (after
Carlgren, 1949
; additions in bold and modifications in italics).
Enthemonae
with elongate body with proximal end physa-shaped, rarely flat. Column sometimes without regional differentiation, sometimes divisible into physa, scapus and scapulus. No sphincter. Tentacles few up to 40, the inner not shorter than the outer ones.
Often with
a single siphonoglyph, rarely with two
indistinct
ones. Perfect pairs of mesenteries variable in number up to 20. Retractors of perfect mesenteries strong, diffuse to restricted, more or less reniform. Parietal muscles distinct. Cnidom: gracile spirocysts, basitrichs, holotrichs,
b-
mastigophores and p-
mastigophores A
.
Type
genus:
Halcampoides
Danielssen, 1890
I n c l u d e d g e n e r a: A c t h e l m i s
L ü t k e n, 1 8 7 5 (?),
Calamactinia
Carlgren, 1949
(?),
Calamactis
Carlgren, 1951
(?),
Halcampella
Andres, 1883
(?),
Pentactinia
Carlgren, 1900
(?),
Scytophorus
Hertwig, 1882
and
Siphonactinopsis
Carlgren, 1921
(?).
Remarks:
Rodríguez
et al
. (2012)
synonymized
Halcampoididae
with
Halcampidae
based on the position of an unidentified species of
Halcampoides
(
type
genus of
Halcampoididae
) as the sister-group to an unidentified species of
Halcampa
Gosse, 1858
(
type
genus of
Halcampidae
). Based on the results of our phylogenetic analyses, we consider
Halcampidae
and
Halcampoididae
as separate families (
Studer, 1879
). Thus, we resurrect family
Halcampoididae
and reclassify it as an actinostoloidean, given its position in our phylogenetic analysis. We include
H. purpureus
and
S. striatus
Hertwig, 1882
until a complete revision of remaining former halcampoidid genera.