Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities Author Brazidec, Manuel https://orcid.org/0000-0002-0860-8972 Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China manuel.brazidec@gmail.com Author Vilhelmsen, Lars https://orcid.org/0000-0002-5593-5722 Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark Author Boudinot, Brendon E. https://orcid.org/0000-0002-4588-0430 Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany Author Richter, Adrian https://orcid.org/0000-0001-5627-2302 Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan Author Hammel, Joerg U. https://orcid.org/0000-0002-6744-6811 Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany Author Perkovsky, Evgeny E. https://orcid.org/0000-0002-7959-4379 Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine Author Fan, Yong Fushun Amber Institute, Fushun 113005, China Author Wang, Zhen Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China Author Wu, Qiong Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China Author Wang, Bo State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China Author Perrichot, Vincent https://orcid.org/0000-0002-7973-0430 Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France text Arthropod Systematics & amp; Phylogeny 2024 2024-03-22 82 151 181 http://dx.doi.org/10.3897/asp.82.e111148 journal article http://dx.doi.org/10.3897/asp.82.e111148 1864-8312-82-151 43AC036E93CC4D79939A07DF54BE1A2D 8D187145BFCA55A1A73960CDE5BD124B 3.1.1.1.1. † Cretolyra noijebumensis sp. nov. Figures 2A , 3 Etymology. The specific epithet is an adjective referring to the type locality where the amber pieces were collected. Material studied. Holotype male IGR.BU-067; housed in the amber collection of the Geology Department and Museum of the University of Rennes , France (IGR). Type locality. Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar. Age. Upper Albian to lower Cenomanian, mid-Cretaceous. Diagnosis. Antennae inserted on edge of clypeus (Fig. 3A ; vs. inserted above clypeal margin in Cretolyra shawi gen. et sp. nov. ); axillae contiguous medially and axillar grooves crenulated (Fig. 3B ; vs axillae separated medially and axillar grooves smooth in Cretolyra shawi gen. et sp. nov. ). Description. Body length 3.33 mm; body very pubescent, especially on metasoma where setae conceal margins of tergites; the setae have agglomerated small bubbles that make observation difficult. - Head globular, higher than long (length 0.55 mm; height 0.75 mm); frons convex, smooth, divided by thin median sulcus; compound eye oval, higher than long; vertex convex, with smooth ocellar triangle; clypeus well-developed; torulus inserted closer to clypeus than to eye; shallow but distinct subantennal groove adjoining ventral margin of eye; antenna half as long as body, almost reaching metasoma; scape twice as long as wide (length 0.16 mm); pedicel shorter than scape (length 0.11 mm); flagellomeres cylindrical, elongate, longer than wide; flagellomere 1 shortest (length 0.15 mm); following flagellomeres longer (length ca. 0.21 mm); occipital carina crenulate. - Mesosoma almost half body length (length 1.29 mm; height 0.65 mm); mesoscutum convex, shagreened, divided by smooth median mesoscutal sulcus (mesoscutum length 0.58 mm; width ~0.60 mm); parapsidal lines present, diverging anteriorly; axillae contiguous medially, axillar groove crenulate; mesoscutellum diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, with incomplete pairs of median and submedian carinae, two complete lateral longitudinal carinae, anterior region of propodeum with transversal row of foveae adjacent to concealed metanotum, posterior region of propodeum with three large foveae. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length 2.50 mm); C, Sc+R, M+Cu, A, Rs fully pigmented; R1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell rectangular, narrow; Cu almost reaching posterior wing margin. - Legs covered with setae; two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus three times as long as following metatarsomeres and with row of short comb-like setae along ventral surface. - Metasoma half body length (length 1.48 mm), elongate and narrowed at apex; seven smooth tergites with pubescent posterior margin; first tergite largest; second tergite one quarter of metasoma length; remaining tergites shorter.