New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil
Author
Khaustov, Alexander A.
Author
Frolov, Andrey V.
text
Zootaxa
2017
4294
5
501
521
journal article
32629
10.11646/zootaxa.4294.5.1
73bbb6a1-db6a-40d3-b056-883d65441665
1175-5326
833794
5E39DFDF-A04D-4EA6-B138-16908DA99F8F
Key to world species of
Pavania
(based on
Hajiqanbar & Khaustov 2010
)
1. Setae
sc
1 absent.......................................................................................2
- Setae
sc
1 present...................................................................................... 5
2. Setae 1
c
and 2
c
present................................................................................. 3
- Setae 1
c
and 2
c
absent...........................................................
P
.
neotropica
sp. nov.
(
Brazil
)
3. Setae
v
1
shorter than distance between their bases; setae
cha
less than three times longer than
chb
; setae
e
never longer than
f
; setae
h
2 at most seven times longer than
h
1..................................................................4
- Setae
v
1
longer than distance between their bases; setae
cha
three times longer than
chb
; setae
e
longer than
f
; setae
h
2 15 times longer than
h
1...............................................
P
.
gymnopleuri
Hajiqanbar and Khaustov, 2010
(
Iran
)
4. Genu I with one seta (
v’
); dorsal idiosomal setae smooth; setae
c
1 longer than
c
2; setae
c
1 and
d
pointed............................................................................
P
.
sabzevarensis
Hajiqanbar and Khaustov, 2010
(
Iran
)
- Genu I with two setae (
v’
,
l’
); dorsal idiosomal setae weakly barbed; setae
c
2 longer than
c
1; setae
c
1 and
d
distinctly blunt- ended.....................................................
P
.
onthophagi
Hajiqanbar and Hajiqanbar, 2010
(
Iran
)
5. Setae
sc
1 capitate......................................................................................6
- Setae
sc
1 seta-like.............................................
P. setiformis
Loghmani and Hajiqanbar, 2013
(Iran)
6. Setae (
u
) and (
pv
) of tarsus I not lanceolate.................................................................7
- Setae (
u
) and (
pv
) of tarsus I lanceolate..........................
P
.
lanceolata
Bahramian and Hajiqanbar, 2015
(Iran)
7. Setae
h
2 less than 3.5 times longer than
h
1..................................................................8
- Setae
h
2 more than 3.5 times longer than
h
1................................................................ 1 5
8. Setae
c
1 never reaching beyond bases of setae
f
; setae
c
1 shorter than
h
2; setae
d
shorter than
h
2............................................. 9
- Setae
c
1 reaching beyond bases of setae
f
; setae
c
1 longer than
h
2; setae
d
and
h
2 subequal....................................................................................................
P
.
perhirsuta
Mahunka, 1973
(
Ghana
)
9. Setae
h
1 longer than or subequal to
d
,
e
and
f...............................................................
10
- Setae
h
1 shorter than
d
,
e
and
f
..........................................................................14
10. Setae
h
1 shorter than
sc
2; setae
sc
2 longer than or subequal to distance between their bases; seta
f
inserted posteriorly to seta
e.
................................................................................................... 11
- Setae
h
1 longer than
sc
2; setae
sc
2 distinctly shorter than distance between their bases; seta
e
inserted at the same level as seta
f......................................................................
P
.
equisetosa
Mahunka, 1975
(
Ghana
)
11. Setae
sc
2 distinctly longer than distance between their bases; setae
sc
2 reaching beyond bases of setae
c
1............................. 12
- Setae
sc
2 subequal to distance between their bases; setae
sc
2 never reaching beyond bases of setae
c
1...................13
12. Setae
c
1 longer than
c
2; setae
h
1 longer than
e
; setae
h
1 and
v
1
subequal......
P
.
riparia
Sevastianov, 1980
(
Ukraine
,
Slovakia
)
- Setae
c
2 longer than
c
1; setae
h
1 and
e
subequal; setae
h
1 longer than
v
1.
................
P
.
luisiae
Mahunka, 1974
(
Ghana
)
13. Setae
c
1,
d
,
e
and
f
blunt-ended; setae
v
1
reaching beyond bases of setae
cha
..........................................................................................
P
.
carabidophila
Khaustov, 2005
(
Russia
:
Krasnodarskiy Kray
)
- Setae
c
1,
d
,
e
and
f
pointed; setae
v
1
never reaching beyond bases of setae
cha........................................................................................................
P
.
bembidii
Khaustov, 2005
(
Russia
:
Crimea
)
14. Setae
h
2 more than twice as long as
h
1; setae
e
and
f
subequal and both longer than
d
; setae
f
distinctly longer than
c
1.........................................................................
P
.
tahanae
Sevastianov and Abo-Korah, 1985
(
Egypt
)
- Setae
h
2 less than twice as long as
h
1; setae
e
and
d
subequal and both longer than
f
; setae
c
1 and
f
subequal.....................................................................
P
.
protracta
Sevastianov, 1980
(Russia, Turkmenistan, Iran)
15. Setae
h
2 more than six times longer than
h
1................................................................. 1 6
- Setae
h
2 less than six times longer than
h
1..................................................................19
16. Setae
sc
2 less than 2.5 times longer than
v
1
; setae
f
less than twice as long as
e
; setae
e
shorter than
v
1.
.................17
- Setae
sc
2 at least 3.5 times longer than
v
1
;
setae
f
more than twice as long as
e
; setae
e
longer than
v
1.
...........................................................................................
P
.
endroedyi
Mahunka, 1975
(
Ghana
)
17. Setae
sc
2 more than twice as long as
v
1
; setae
f
and
d
subequal; setae
c
1 never reaching beyond posterior border of tergite C......................................................................................................18
- Setae
sc
2 less than twice as long as
v
1
; setae
f
longer than
d
; setae
c
1 reaching beyond posterior border of tergite C................................................................................
P
.
brasiliensis
Mahunka, 1970 (
Brazil
)
18. Setae 2
a
as long as 2
c
and both longer than
c
1,
d
and
f
; setae
m
protruding beyond anterior border of gnathosoma........................................................................
P
.
elongata
Hajiqanbar and Khaustov, 2010
(
Iran
)
- Setae 2
a
longer than 2
c
and both shorter than
c
1,
d
and
f
; setae
m
never protruding beyond anterior border of gnathosoma.............................................................................
P
.
simplex
Mahunka, 1973
(
Ghana
)
19. Setae
f
more than twice as long as
e
; setae
e
and
h
1 subequal; setae on coxal fields II not subequal.....................20
- Setae
e
and
f
subequal; setae
e
longer than
h
1; setae on coxal fields II subequal................................................................................................
P
.
tadjikistanica
Sevastianov, 1980
(Tadjikistan, Iran)
20. Most dorsal idiosomal setae weakly barbed and blunt-ended; setae
c
1 longer than
c
2; setae
sc
2 less than twice as long as
c
1..................................................................
P
.
kamalii
Hajiqanbar and Khaustov, 2010
(
Iran
)
- Dorsal idiosomal setae smooth and pointed; setae
c
2 longer than
c
1; setae
sc
2 more than twice as long as
c
1..................................................................................
P
.
fusiformis
Lombardini, 1949
(Italy, Iran)