A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215 Author Van Bakel, Barry W. M. Author Guinot, Danièle Author Artal, Pedro Author Fraaije, René H. B. Author Jagt, John W. M. text Zootaxa 2012 2012-02-29 3215 1 1 216 https://biotaxa.org/Zootaxa/article/view/zootaxa.3215.1.1 journal article 10.11646/zootaxa.3215.1.1 1175­5334 5248640 B20CD4A6-D150-4CCF-931F-ED6D7EA54E8C Genus Heus Bishop & Williams, 2000 Heus Bishop & Williams, 2000: 290 . Type species. Heus foersteri Bishop & Williams, 2000 . Remarks. Only the carapace of Heus foersteri (see Bishop & Williams 2000: 290 , fig. 4), from the upper Campanian of South Dakota and Colorado , U.S.A. , is preserved. This reveals a unique set of characters: rostrum broad and narrow, strongly sulcate, with a keel that extending onto the carapace; oval swellings on epigastric and protogastric regions; cervical groove discernible; lateral margins with but a single, diminutive spine. These characters distinguish it easily from Bournelyreidus n. gen. Re-examination of the type series of B. ? oaheensis (see above) is needed to decide if it might also be better accommodated in Heus . The wide carapace, with clearly concave posterior (divergent) portions of the posterolateral margins, weakly armoured anterolateral margin, and swellings on the anterior carapace surface are somewhat similar to the configuration in Symethis Weber, 1795 (see below). The orbitofrontal construction, however, makes its placement in the Lyreidinae more likely.