Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini) Author Perger, Robert Author Grossi, Paschoal Coelho text ZooKeys 2013 346 1 16 http://dx.doi.org/10.3897/zookeys.346.6114 journal article http://dx.doi.org/10.3897/zookeys.346.6114 1313-2970-346-1 Oryctophileurus nasicornis (Burmeister, 1847) Fig. 3 Material examined. Colombia: Cauca dep., Cauca Valley: 1 male with body length 19 mm, det. Endroedi 1976 (ZMHB); Boyaca dep., Muzo muni.: 1 female with body length 20 mm, S. Apollin coll., det. Ohaus, revised by Endroedi (1977) (ZMHB). Diagnosis. Color dark brown, moderately shining. Horn on frons not projecting above anterior edges of pronotal protuberance, weakly recurved, in female with truncate apex; ocular canthus rounded in male, and subquadrate in female. Pronotum broadly rounded, more in female. Distance between inner teeth of dorsal pronotal protuberance narrower than distance between eyes. Pronotal protuberance on same level with elytra. Development of horn and pronotal protuberance equal in both sexes. Elytra striate, with five well defined discal striae; striae regular, at sides weakly defined; punctures regular, about the same size of those at disc. Meso and metatibiae with 4 to 5 distal teeth. Pygidium regularly convex in both sexes, more in female, and densely and finely wrinkled. Parameres with basal half broad, apex slender, straight. Figure 3. Dorsal and lateral habitus of Oryctophileurus nasicornis (Burmeister, 1847) A and B male C and D female, scale bar 5 mm. Geographical distribution. The species is known only from the Cauca Valley in Colombia, which is surrounded by the parallel, peninsula-like projections of the Andean Cordillera Occidental and Cordillera Central and the Muzo municipality which is situated on the eastern slope of the Cordillera Oriental (Fig. 1A). These areas include wet premontane forest and, to a lesser extent, upper montane forests and paramos ( Espinal 1992 ).