Costal vein chaetotaxy, a neglected character source in Fanniidae and Muscidae (Diptera: Calyptratae) Author Michelsen, Verner 9BD4846E-F4D0-4DB2-A567-FAF0A58B6D98 Natural History Museum of Denmark, University of Copenhagen, Denmark. vmichelsen@snm.ku.dk text European Journal of Taxonomy 2022 2022-06-28 826 94 134 http://dx.doi.org/10.5852/ejt.2022.826.1839 journal article 86210 10.5852/ejt.2022.826.1839 ed37ce13-a67c-4146-b99e-db8ede0a145b 2118-9773 6784127 34BA1AB7-6107-4636-9645-B1C0216DCE5E Subfamily Reinwardtiinae Brauer & Bergenstamm, 1889 This taxon is a heterogeneous, para- or polyphyletic assemblage of genera which is also reflected in its diverse costal chaetotaxy. Species from seven among the 15 currently recognized genera ( Table 1 ) were examined. • Muscina Robineau-Desvoidy, 1830 ( Fig. 12A ), Passeromyia Rodhain & Villeneuve, 1915 , Philornis Meinert, 1890 The examined species belonging to the above genera of Reinwardtiinae share the prevalent muscid state A 4 in which vein C is bare dorsally and extensively setulose ventrally: Muscina (5 spp), Passeromyia heterochaeta Villeneuve, 1915 and P. steini Pont, 1970 , Philornis (5 spp). Fig. 11. Atherigona orientalis Schiner, 1868 , ♀. A . Wing, ventral view. B . Detail of wing showing CS1‒3, ventral view. • Synthesiomyia Brauer & Bergenstamm, 1893 The only included species, Synthesiomyia nudipes , has vein C only fully setulose ventrally on CS2 (state A2 ), although some irregular ventral costal setulae extent onto CS2 or (females only) even the basal part of CS3. • Brachygasterina Macquart, 1851 , Palpibracus Rondani, 1863 ( Fig. 12B ), Psilochaeta Stein, 1911 The examined species of these three exclusively South American genera differ substantially from the preceding reinwardtiine genera by the dorsally extensively setulose vein C (state A8 , Fig. 12B ): Brachygasterina fulvohumeralis (Malloch, 1922) , B. violaceiventris Macquart, 1851 , Palpibracus (12 spp) and Psilochaeta chalybea (Wiedemann, 1830) . Only Brachygasterina major Malloch, 1934 differs by having the dorsal setulae restricted to CS2 and CS3 (females) or CS3 (males) (state A7 ).